Finding intermodel H-bonds Finding intramodel H-bonds Constraints relaxed by 0.4 angstroms and 20 degrees Models used: #0 4mvj_G.cif H-bonds (donor, acceptor, hydrogen, D..A dist, D-H..A dist): THR 2.A OG1 GLU 28.A OE2 no hydrogen 3.232 N/A ILE 3.A N ASP 26.A O no hydrogen 2.920 N/A LYS 4.A N ASP 90.A OD2 no hydrogen 3.143 N/A LYS 4.A NZ THR 2.A O no hydrogen 3.463 N/A LYS 4.A NZ THR 2.A OG1 no hydrogen 3.130 N/A VAL 5.A N GLU 28.A O no hydrogen 2.822 N/A GLY 6.A N VAL 91.A O no hydrogen 2.996 N/A ILE 7.A N ALA 31.A O no hydrogen 2.934 N/A ASN 8.A N ALA 93.A O no hydrogen 2.990 N/A ASN 8.A ND2 GLU 94.A OE1 no hydrogen 2.905 N/A GLY 9.A N ASN 33.A O no hydrogen 2.868 N/A PHE 10.A N ASP 34.A OD2 no hydrogen 3.005 N/A ARG 12.A NE GLU 313.A OE1 no hydrogen 2.789 N/A ARG 12.A NH1 ASP 47.A OD2 no hydrogen 2.884 N/A ARG 12.A NH2 ASP 47.A OD1 no hydrogen 3.240 N/A ARG 12.A NH2 ASP 47.A OD2 no hydrogen 3.282 N/A ARG 12.A NH2 GLU 313.A OE1 no hydrogen 3.304 N/A ARG 12.A NH2 GLU 313.A OE2 no hydrogen 2.910 N/A ARG 15.A N GLY 11.A O no hydrogen 2.901 N/A ARG 15.A NH1 PHE 10.A O no hydrogen 3.062 N/A ILE 16.A N ARG 12.A O no hydrogen 3.078 N/A VAL 17.A N ILE 13.A O no hydrogen 2.813 N/A PHE 18.A N GLY 14.A O no hydrogen 2.929 N/A ARG 19.A N ARG 15.A O no hydrogen 3.067 N/A ARG 19.A NH1 HIS 50.A O no hydrogen 3.312 N/A ARG 19.A NH1 GLY 51.A O no hydrogen 2.971 N/A ARG 19.A NH2 LEU 45.A O no hydrogen 2.852 N/A ARG 19.A NH2 GLY 51.A O no hydrogen 3.007 N/A ALA 20.A N ILE 16.A O no hydrogen 2.905 N/A ALA 21.A N VAL 17.A O no hydrogen 2.984 N/A GLN 22.A N ARG 19.A O no hydrogen 3.241 N/A GLN 22.A NE2 PHE 18.A O no hydrogen 3.334 N/A LYS 23.A N ALA 20.A O no hydrogen 3.226 N/A ARG 24.A N ALA 21.A O no hydrogen 3.300 N/A ARG 24.A NE ASP 26.A OD1 no hydrogen 3.166 N/A ARG 24.A NH1 ASP 322.A OD1 no hydrogen 2.815 N/A ARG 24.A NH2 ASP 26.A OD2 no hydrogen 3.101 N/A GLU 28.A N ILE 3.A O no hydrogen 3.057 N/A VAL 30.A N VAL 5.A O no hydrogen 2.956 N/A ILE 32.A N ARG 72.A O no hydrogen 2.794 N/A ASN 33.A N ILE 7.A O no hydrogen 2.831 N/A ASN 33.A ND2 ASN 8.A OD1 no hydrogen 2.806 N/A ASN 33.A ND2 GLU 76.A O no hydrogen 3.010 N/A ASP 34.A N THR 74.A O no hydrogen 3.188 N/A LEU 35.A N ASP 34.A OD1 no hydrogen 2.649 N/A TYR 40.A N ASP 37.A O no hydrogen 3.112 N/A MET 41.A N ASP 37.A O no hydrogen 3.025 N/A ALA 42.A N ALA 38.A O no hydrogen 2.934 N/A TYR 43.A N ASP 39.A O no hydrogen 3.264 N/A MET 44.A N TYR 40.A O no hydrogen 2.935 N/A LEU 45.A N MET 41.A O no hydrogen 2.948 N/A LEU 45.A N ALA 42.A O no hydrogen 3.213 N/A THR 49.A N ASP 47.A OD1 no hydrogen 2.879 N/A HIS 50.A N ASP 47.A OD1 no hydrogen 2.857 N/A HIS 50.A NE2 GLU 313.A OE2 no hydrogen 2.835 N/A GLY 55.A N PHE 53.A O no hydrogen 3.027 N/A THR 56.A N ASN 67.A OD1 no hydrogen 3.000 N/A GLU 58.A N ILE 65.A O no hydrogen 2.928 N/A LYS 60.A N HIS 63.A O no hydrogen 2.859 N/A HIS 63.A N LYS 60.A O no hydrogen 3.008 N/A LEU 64.A N ILE 71.A O no hydrogen 3.298 N/A ILE 65.A N GLU 58.A O no hydrogen 2.774 N/A VAL 66.A N LYS 69.A O no hydrogen 3.068 N/A ASN 67.A N THR 56.A O no hydrogen 2.861 N/A ASN 67.A ND2 GLN 22.A OE1 no hydrogen 3.339 N/A LYS 69.A N VAL 66.A O no hydrogen 3.174 N/A ILE 71.A N LEU 64.A O no hydrogen 2.836 N/A ARG 72.A N VAL 30.A O no hydrogen 3.036 N/A ARG 72.A NH1 LEU 82.A O no hydrogen 3.230 N/A ARG 72.A NH1 LYS 83.A O no hydrogen 2.817 N/A ARG 72.A NH2 LEU 82.A O no hydrogen 2.937 N/A VAL 73.A N HIS 63.A ND1 no hydrogen 3.156 N/A THR 74.A N ILE 32.A O no hydrogen 3.115 N/A THR 74.A OG1 ILE 32.A O no hydrogen 3.531 N/A THR 74.A OG1 ASN 33.A OD1 no hydrogen 2.847 N/A GLU 76.A N ASN 33.A OD1 no hydrogen 2.934 N/A ALA 80.A N ASP 78.A OD1 no hydrogen 3.531 N/A ASN 81.A N ASP 78.A O no hydrogen 3.050 N/A LEU 82.A N PRO 79.A O no hydrogen 3.205 N/A TRP 84.A N ALA 111.A O no hydrogen 2.806 N/A TRP 84.A NE1 HIS 108.A ND1 no hydrogen 2.996 N/A GLU 86.A N LYS 83.A O no hydrogen 3.228 N/A VAL 87.A N TRP 84.A O no hydrogen 3.164 N/A GLY 88.A N ASP 85.A O no hydrogen 3.145 N/A VAL 89.A N TRP 84.A O no hydrogen 3.178 N/A ASP 90.A N LYS 4.A O no hydrogen 2.828 N/A VAL 91.A N LYS 4.A O no hydrogen 3.175 N/A VAL 92.A N LYS 115.A O no hydrogen 2.914 N/A ALA 93.A N GLY 6.A O no hydrogen 2.821 N/A GLU 94.A N VAL 117.A O no hydrogen 2.937 N/A ALA 95.A N ASN 8.A O no hydrogen 2.683 N/A THR 96.A N GLU 94.A OE2 no hydrogen 2.994 N/A THR 96.A OG1 GLU 94.A OE2 no hydrogen 2.513 N/A LEU 98.A N THR 96.A OG1 no hydrogen 3.020 N/A THR 101.A N THR 104.A OG1 no hydrogen 3.241 N/A THR 101.A OG1 THR 104.A OG1 no hydrogen 2.921 N/A ASP 102.A N ASP 124.A OD2 no hydrogen 2.604 N/A THR 104.A N THR 101.A OG1 no hydrogen 3.413 N/A THR 104.A OG1 THR 101.A O no hydrogen 3.045 N/A THR 104.A OG1 THR 101.A OG1 no hydrogen 2.921 N/A ALA 105.A N THR 101.A O no hydrogen 3.058 N/A ARG 106.A N ASP 102.A O no hydrogen 2.777 N/A ARG 106.A NE ASP 102.A OD2 no hydrogen 3.471 N/A ARG 106.A NH1 ASP 142.A OD1 no hydrogen 3.296 N/A ARG 106.A NH1 ASP 142.A OD2 no hydrogen 2.853 N/A ARG 106.A NH2 ASP 102.A OD1 no hydrogen 2.693 N/A ARG 106.A NH2 ASP 102.A OD2 no hydrogen 3.285 N/A ARG 106.A NH2 ASP 142.A OD1 no hydrogen 3.538 N/A HIS 108.A N ALA 105.A O no hydrogen 3.028 N/A HIS 108.A NE2 GLU 94.A OE1 no hydrogen 2.905 N/A ILE 109.A N ARG 106.A O no hydrogen 2.841 N/A THR 110.A N ARG 106.A O no hydrogen 3.055 N/A THR 110.A OG1 ARG 106.A O no hydrogen 3.501 N/A ALA 111.A N LYS 107.A O no hydrogen 2.883 N/A GLY 112.A N ILE 109.A O no hydrogen 3.028 N/A ALA 113.A N HIS 108.A O no hydrogen 2.994 N/A LYS 114.A N ASP 90.A O no hydrogen 2.953 N/A LYS 115.A N ASP 90.A O no hydrogen 3.084 N/A LYS 115.A NZ TYR 138.A OH no hydrogen 2.997 N/A LYS 115.A NZ ILE 327.A O no hydrogen 2.741 N/A VAL 116.A N ASP 142.A O no hydrogen 2.859 N/A VAL 117.A N VAL 92.A O no hydrogen 2.857 N/A MET 118.A N VAL 144.A O no hydrogen 2.768 N/A THR 119.A N GLU 94.A O no hydrogen 2.977 N/A THR 119.A OG1 GLU 94.A O no hydrogen 3.116 N/A SER 122.A OG ASP 124.A O no hydrogen 3.569 N/A SER 122.A OG THR 126.A O no hydrogen 2.661 N/A LYS 123.A N LEU 100.A O no hydrogen 2.866 N/A THR 126.A N ASP 124.A OD1 no hydrogen 2.870 N/A THR 126.A OG1 ASP 124.A OD1 no hydrogen 2.625 N/A THR 126.A OG1 ASP 124.A OD2 no hydrogen 3.402 N/A PHE 129.A N SER 145.A O no hydrogen 2.818 N/A VAL 130.A N ASN 134.A OD1 no hydrogen 2.767 N/A ALA 133.A N VAL 130.A O no hydrogen 2.885 N/A ASN 134.A N VAL 130.A O no hydrogen 3.071 N/A ASN 134.A ND2 MET 128.A O no hydrogen 3.037 N/A LYS 137.A N ASN 134.A O no hydrogen 3.003 N/A TYR 138.A N PHE 135.A O no hydrogen 2.923 N/A TYR 138.A OH GLN 141.A O no hydrogen 2.459 N/A ALA 139.A N GLN 141.A OE1 no hydrogen 2.942 N/A GLN 141.A N GLN 141.A OE1 no hydrogen 2.624 N/A VAL 144.A N VAL 116.A O no hydrogen 2.980 N/A SER 145.A N PRO 127.A O no hydrogen 2.983 N/A SER 145.A OG GLY 120.A O no hydrogen 2.430 N/A ASN 146.A N MET 118.A O no hydrogen 3.099 N/A ASN 146.A ND2 PHE 129.A O no hydrogen 3.161 N/A ALA 147.A N SER 145.A OG no hydrogen 3.143 N/A SER 148.A OG THR 150.A OG1 no hydrogen 2.896 N/A THR 149.A OG1 TYR 310.A OH no hydrogen 2.612 N/A THR 150.A N SER 148.A OG no hydrogen 3.145 N/A THR 150.A OG1 SER 148.A OG no hydrogen 2.896 N/A THR 150.A OG1 ALA 209.A O no hydrogen 3.163 N/A ASN 151.A N SER 148.A O no hydrogen 3.025 N/A ASN 151.A ND2 ASN 146.A O no hydrogen 3.056 N/A ASN 151.A ND2 SER 148.A O no hydrogen 2.796 N/A CYS 152.A N THR 149.A O no hydrogen 3.008 N/A CYS 152.A SG SER 289.A OG no hydrogen 3.622 N/A CYS 152.A SG SER 308.A O no hydrogen 3.562 N/A CYS 152.A SG SER 308.A OG no hydrogen 3.612 N/A LEU 153.A N THR 149.A O no hydrogen 2.958 N/A ALA 154.A N THR 150.A O no hydrogen 2.931 N/A ALA 157.A N LEU 153.A O no hydrogen 2.887 N/A LYS 158.A N ALA 154.A O no hydrogen 2.844 N/A LYS 158.A NZ ASP 162.A OD2 no hydrogen 2.908 N/A VAL 159.A N PRO 155.A O no hydrogen 3.234 N/A ILE 160.A N LEU 156.A O no hydrogen 2.994 N/A ASN 161.A N ALA 157.A O no hydrogen 2.751 N/A ASP 162.A N LYS 158.A O no hydrogen 2.809 N/A ASN 163.A N VAL 159.A O no hydrogen 3.048 N/A ASN 163.A ND2 VAL 159.A O no hydrogen 2.915 N/A PHE 164.A N ILE 160.A O no hydrogen 2.752 N/A GLY 165.A N ASN 161.A O no hydrogen 2.774 N/A ILE 166.A N ASN 161.A OD1 no hydrogen 2.795 N/A ILE 167.A N ARG 244.A O no hydrogen 2.909 N/A GLY 169.A N LYS 223.A O no hydrogen 2.823 N/A LEU 170.A N THR 242.A O no hydrogen 2.946 N/A MET 171.A N THR 225.A O no hydrogen 3.023 N/A THR 172.A N ASP 240.A O no hydrogen 2.966 N/A THR 173.A N MET 227.A O no hydrogen 2.983 N/A THR 173.A OG1 VAL 238.A O no hydrogen 3.207 N/A THR 173.A OG1 TYR 310.A OH no hydrogen 2.921 N/A VAL 174.A N VAL 238.A O no hydrogen 2.936 N/A HIS 175.A N PHE 229.A O no hydrogen 2.778 N/A HIS 175.A ND1 ALA 176.A O no hydrogen 2.782 N/A ALA 176.A N SER 237.A OG no hydrogen 3.109 N/A THR 177.A N THR 233.A O no hydrogen 2.773 N/A THR 177.A OG1 VAL 231.A O no hydrogen 3.019 N/A THR 178.A N GLN 181.A OE1 no hydrogen 3.265 N/A THR 180.A OG1 THR 178.A OG1 no hydrogen 3.292 N/A GLN 181.A N THR 178.A O no hydrogen 3.065 N/A GLY 186.A N GLY 195.A O no hydrogen 2.823 N/A SER 188.A OG ASP 191.A O no hydrogen 2.875 N/A ASP 191.A N SER 188.A O no hydrogen 3.074 N/A GLY 194.A N ASP 191.A OD1 no hydrogen 3.375 N/A GLY 195.A N TRP 192.A O no hydrogen 3.121 N/A ARG 196.A N ARG 193.A O no hydrogen 3.077 N/A ARG 196.A NE ASP 185.A OD1 no hydrogen 3.287 N/A ARG 196.A NE ASP 185.A OD2 no hydrogen 3.185 N/A GLY 197.A N ASP 185.A OD1 no hydrogen 2.754 N/A ALA 198.A N LYS 182.A O no hydrogen 2.871 N/A ASN 201.A ND2 GLY 197.A O no hydrogen 2.900 N/A ILE 203.A N ARG 230.A O no hydrogen 2.811 N/A SER 205.A N ALA 228.A O no hydrogen 2.703 N/A SER 205.A OG SER 206.A O no hydrogen 3.196 N/A ALA 212.A N GLY 208.A O no hydrogen 2.661 N/A VAL 213.A N ALA 210.A O no hydrogen 3.349 N/A GLY 214.A N LYS 211.A O no hydrogen 2.897 N/A LYS 215.A N ALA 212.A O no hydrogen 3.017 N/A LEU 217.A N VAL 213.A O no hydrogen 2.729 N/A GLU 219.A N GLU 219.A OE1 no hydrogen 2.625 N/A LEU 220.A N LEU 217.A O no hydrogen 2.782 N/A ASN 221.A N PRO 218.A O no hydrogen 2.872 N/A LYS 223.A N LEU 220.A O no hydrogen 3.045 N/A MET 227.A N MET 171.A O no hydrogen 2.990 N/A ALA 228.A N SER 205.A O no hydrogen 2.928 N/A PHE 229.A N THR 173.A O no hydrogen 2.908 N/A ARG 230.A N ILE 203.A O no hydrogen 2.790 N/A ARG 230.A NH1 THR 178.A OG1 no hydrogen 2.977 N/A ARG 230.A NH1 GLN 181.A OE1 no hydrogen 2.735 N/A VAL 231.A N HIS 175.A O no hydrogen 2.908 N/A THR 233.A N VAL 231.A O no hydrogen 2.874 N/A ASN 235.A ND2 ASN 283.A OD1 no hydrogen 2.773 N/A ASN 235.A ND2 ASP 311.A OD1 no hydrogen 2.898 N/A ASN 235.A ND2 ASP 311.A OD2 no hydrogen 3.373 N/A VAL 236.A N ASN 312.A OD1 no hydrogen 2.856 N/A SER 237.A N TYR 310.A O no hydrogen 2.843 N/A SER 237.A OG ASN 312.A OD1 no hydrogen 2.935 N/A VAL 238.A N VAL 174.A O no hydrogen 2.971 N/A VAL 239.A N SER 308.A O no hydrogen 2.769 N/A ASP 240.A N THR 172.A O no hydrogen 2.701 N/A LEU 241.A N LEU 306.A O no hydrogen 2.787 N/A THR 242.A N LEU 170.A O no hydrogen 2.912 N/A VAL 243.A N VAL 304.A O no hydrogen 2.846 N/A ARG 244.A N GLU 168.A O no hydrogen 2.976 N/A ARG 244.A NH1 GLU 168.A OE2 no hydrogen 2.582 N/A LEU 245.A N ASN 302.A O no hydrogen 2.898 N/A GLU 246.A N GLY 165.A O no hydrogen 2.805 N/A LYS 247.A N PHE 164.A O no hydrogen 2.865 N/A ALA 249.A N ASP 301.A O no hydrogen 2.852 N/A THR 250.A N GLN 253.A OE1 no hydrogen 2.841 N/A THR 250.A OG1 GLN 253.A OE1 no hydrogen 3.304 N/A TYR 251.A OH ALA 293.A O no hydrogen 2.602 N/A GLN 253.A N THR 250.A OG1 no hydrogen 3.228 N/A ILE 254.A N THR 250.A O no hydrogen 3.208 N/A LYS 255.A N TYR 251.A O no hydrogen 2.934 N/A LYS 255.A NZ GLU 252.A OE2 no hydrogen 3.440 N/A ALA 256.A N GLU 252.A O no hydrogen 3.082 N/A ALA 257.A N GLN 253.A O no hydrogen 2.962 N/A VAL 258.A N ILE 254.A O no hydrogen 3.037 N/A LYS 259.A N LYS 255.A O no hydrogen 2.874 N/A ALA 260.A N ALA 256.A O no hydrogen 2.919 N/A ALA 261.A N ALA 257.A O no hydrogen 3.109 N/A ALA 262.A N VAL 258.A O no hydrogen 2.801 N/A GLU 263.A N LYS 259.A O no hydrogen 3.056 N/A GLY 264.A N ALA 261.A O no hydrogen 3.255 N/A MET 266.A N ALA 261.A O no hydrogen 2.953 N/A LYS 267.A N GLY 264.A O no hydrogen 3.151 N/A VAL 269.A N MET 266.A O no hydrogen 3.149 N/A LEU 270.A N MET 266.A O no hydrogen 3.047 N/A GLY 271.A N SER 289.A O no hydrogen 2.678 N/A TYR 272.A OH GLU 274.A OE2 no hydrogen 2.661 N/A THR 273.A N PHE 291.A O no hydrogen 2.989 N/A ASP 275.A N THR 273.A OG1 no hydrogen 3.107 N/A VAL 278.A N ASP 281.A OD2 no hydrogen 2.958 N/A ASP 281.A N VAL 278.A O no hydrogen 2.881 N/A PHE 282.A N SER 279.A O no hydrogen 2.875 N/A ASN 283.A N THR 280.A O no hydrogen 3.161 N/A ASN 283.A ND2 THR 49.A O no hydrogen 3.111 N/A GLY 284.A N ASP 311.A OD2 no hydrogen 2.692 N/A GLU 285.A N PHE 282.A O no hydrogen 2.872 N/A CYS 287.A N GLU 285.A OE2 no hydrogen 3.196 N/A THR 288.A N VAL 269.A O no hydrogen 2.992 N/A THR 288.A OG1 ASN 151.A O no hydrogen 3.522 N/A THR 288.A OG1 VAL 269.A O no hydrogen 3.496 N/A SER 289.A N VAL 269.A O no hydrogen 3.256 N/A SER 289.A OG CYS 152.A O no hydrogen 2.865 N/A SER 289.A OG SER 308.A OG no hydrogen 2.596 N/A VAL 290.A N TRP 309.A O no hydrogen 2.888 N/A PHE 291.A N GLY 271.A O no hydrogen 2.914 N/A ASP 292.A N VAL 307.A O no hydrogen 2.977 N/A ALA 293.A N THR 273.A O no hydrogen 3.053 N/A LYS 294.A NZ ASP 276.A OD1 no hydrogen 3.530 N/A ALA 295.A N ASP 292.A OD1 no hydrogen 3.065 N/A ILE 297.A N LYS 305.A O no hydrogen 3.056 N/A LEU 299.A N PHE 303.A O no hydrogen 3.027 N/A ASN 300.A N PHE 303.A O no hydrogen 3.404 N/A ASN 302.A N ASN 300.A OD1 no hydrogen 2.773 N/A ASN 302.A ND2 LEU 245.A O no hydrogen 3.099 N/A PHE 303.A N ASN 300.A O no hydrogen 3.348 N/A VAL 304.A N VAL 243.A O no hydrogen 3.035 N/A LYS 305.A N ILE 297.A O no hydrogen 2.916 N/A LYS 305.A NZ ASP 240.A OD1 no hydrogen 3.144 N/A LYS 305.A NZ ASP 240.A OD2 no hydrogen 3.549 N/A LYS 305.A NZ THR 242.A OG1 no hydrogen 2.775 N/A LEU 306.A N LEU 241.A O no hydrogen 2.659 N/A SER 308.A N VAL 239.A O no hydrogen 3.037 N/A SER 308.A OG SER 289.A OG no hydrogen 2.596 N/A TRP 309.A N VAL 290.A O no hydrogen 2.947 N/A TRP 309.A NE1 ASP 292.A OD2 no hydrogen 2.889 N/A TYR 310.A N SER 237.A O no hydrogen 2.894 N/A TYR 310.A OH THR 149.A OG1 no hydrogen 2.612 N/A TYR 310.A OH THR 173.A OG1 no hydrogen 2.921 N/A GLU 313.A N ASP 311.A OD1 no hydrogen 2.913 N/A THR 314.A N ASP 311.A OD1 no hydrogen 3.015 N/A THR 314.A OG1 ASP 311.A OD2 no hydrogen 2.843 N/A GLY 315.A N ASP 311.A O no hydrogen 3.045 N/A TYR 316.A OH ALA 147.A O no hydrogen 2.489 N/A SER 317.A N GLU 313.A O no hydrogen 3.112 N/A SER 317.A OG GLU 313.A O no hydrogen 2.725 N/A ASN 318.A N THR 314.A O no hydrogen 3.063 N/A ASN 318.A ND2 THR 314.A O no hydrogen 2.707 N/A LYS 319.A N GLY 315.A O no hydrogen 2.992 N/A LYS 319.A NZ GLY 268.A O no hydrogen 2.999 N/A LYS 319.A NZ VAL 286.A O no hydrogen 2.937 N/A VAL 320.A N TYR 316.A O no hydrogen 2.891 N/A VAL 320.A N SER 317.A O no hydrogen 3.147 N/A LEU 321.A N SER 317.A O no hydrogen 3.349 N/A ASP 322.A N ASN 318.A O no hydrogen 2.941 N/A LEU 323.A N LYS 319.A O no hydrogen 2.863 N/A ILE 324.A N VAL 320.A O no hydrogen 2.913 N/A ALA 325.A N LEU 321.A O no hydrogen 2.929 N/A HIS 326.A N ASP 322.A O no hydrogen 2.922 N/A ILE 327.A N LEU 323.A O no hydrogen 2.956 N/A SER 328.A N ALA 325.A O no hydrogen 3.277 N/A SER 328.A OG ILE 324.A O no hydrogen 2.806 N/A LYS 329.A N HIS 326.A O no hydrogen 3.356 N/A