1YH8A

Crystal structure of aquifex aeolicus lpxc deacetylase complexed with palmitate
Total Genus 86
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The genus trace: a function that shows values of genus (vertical axis) for subchains spanned between the first residue, and all other residues (shown on horizontal axis). The number of the latter residue and the genus of a given subchain are shown interactively.

Total Genus
86
sequence length
266
structure length
266
Chain Sequence
GLEKTVKEKLSFEGVGIHTGEYSKLIIHPEKEGTGIRFFKNGVYIPARHEFVVHTNHSTDLGFKGQRIKTVEHILSVLHLLEITNVTIEVIGNEIPILDGSGWEFYEAIRKNILNQNREIDYFVVEEPIIVEDEGRLIKAEPSDTLEVTYEGEFKNFLGRQKFTFVEGNEEEIVLARTFAFDWEIEHIKKVGLGKGGSLKNTLVLGKDKVYNPEGLRYENEPVRHKVFDLIGDLYLLGSPVKGKFYSFRGGHSLNVKLVKELAKKQ
Loading diagram...

The genus matrix. At position (x,y) a genus value for a subchain spanned between x’th and y’th residue is shown. Values of the genus are represented by color, according to the scale given on the right.

Structure visualization

After clicking on a point (x,y) in the genus matrix above, a subchain from x to y is shown in color.

Chord Diagram
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publication title UDP-3-O-((R)-3-hydroxymyristoyl)-N-acetylglucosamine Deacetylase Functions through a General Acid-Base Catalyst Pair Mechanism
pubmed doi rcsb
source organism Aquifex aeolicus
molecule tags Hydrolase
molecule keywords UDP-3-O-[3-hydroxymyristoyl] N-acetylglucosamine deacetylase
total genus 86
structure length 266
sequence length 266
chains with identical sequence B
ec nomenclature ec 3.5.1.108: UDP-3-O-acyl-N-acetylglucosamine deacetylase.
pdb deposition date 2005-01-07

pfam database annotations

chain Pfam Accession Code Pfam Family Identifier Pfam Description
A PF03331 LpxC UDP-3-O-acyl N-acetylglycosamine deacetylase
Image from the rcsb pdb (www.rcsb.org)
cath code
ClassArchitectureTopologyHomologyDomain
3.30.230.20 Alpha Beta 2-Layer Sandwich Ribosomal Protein S5; domain 2 lpxc deacetylase, domain 1 1yh8A01
3.30.1700.10 Alpha Beta 2-Layer Sandwich lpxc deacetylase, domain 1 lpxc deacetylase, domain 2 1yh8A02
3P76A 4U3DA 4J3DA 2JT2A 3PS1A 2O3ZA 1YHCA 3P3CA 2IERA 4LCGA 5DRPA 4FW6A 2IESA 1P42A 1XXEA 4IS9A 4OZEA 4U3BA 5DROA 2J65A 3PS2A 4LCHA 2GO3A 4OKGA 4ISAA 5DRQA 4LCFA 4FW4A 3P3GA 3PS3A 5DRRA 4MDTA 2GO4A 1YH8A 3U1YA 4FW3A 4FW7A 4MQYA 3NZKA 4FW5A 3P3EA 3UHMA 2VESA
chains in the Genus database with same CATH superfamily
1Y6ZA 3J7AM 3HKMA 2PO1B 2JE6B 1WUUA 2BA1G 2GS8A 5LMUI 1H74A 1OYPA 2A2DA 2V2QA 4DR5I 4DV5I 1D6TA 4KQVA 4DV2I 4EMDA 3DD6A 1PVGA 2UU9E 4X62I 2BM0A 1U2RA 4B3MI 4JYAI 1HNWE 4JI7E 2WP8A 4DV1I 4DUZI 1IBME 1N34I 5DNLA 4LF5I 3Q6NA 1YHCA 1PKPA 4PRXA 2O1WA 2DEIA 1N34E 2HKJA 2WW4A 1XHKA 2HK3A 4DXLA 2J65A 4WUBA 2Z5CB 2X7IA 4B3ME 3WU5A 3PYDA 3J7AG 5GMDA 1FNMA 3K85A 1J5EE 2O1TA 4AQYE 4QNKA 4FW7A 5C0XB 4DV5E 1K47A 3QT8A 2C39B 1Z5CA 1Z0EA 5BR8I 1MG7A 3P3CA 2E5LI 2UXCI 4IS9A 1IBLI 1N33E 5FLXC 3ZZUA 5LMUE 4USMA 2BV3A 4FW3A 3QT5A 2V2ZA 4FW5A 5LMNI 3PRYA 1Z0CA 1OYSA 3CDIA 3CDJA 1FI4A 1UEKA 4MU3A 2O3ZA 5JEAD 4DR7I 3HJCA 4LF4I 5IWAI 2IESA 3GONA 4LF5E 2UXBI 3LPSA 4OZEA 4DUZE 3F0NA 4DV7I 4NXMI 4HY1A 4JI7I 3PS2A 4LFBE 4X62E 1HNWI 3L7ZB 2WNRB 5A2QQ 4IFDA 2X36A 3ZM7A 1N0VC 4DV2E 3NZKA 5A2QC 3Q1QA 2BM1A 4HXZA 1HNZI 2O1UA 1FWKA 2HHHI 1Y60A 2CG9A 2VQFI 2JEBB 4OX9I 4GQUA 4DR6I 1EA6A 4X66I 2Z5EA 2WNRA 2PO2B 3WU4A 4B3TE 1FJGI 4FWGA 4KHPE 4LCHA 4HACB 4BA2B 4GKKI 4UT4A 2DEJA 2Z5BB 4JI4I 2ZM6E 4LF8I 1Y4SA 4K0KE 2HKEA 4DUYE 4X66E 4DV4I 3J81Q 4DPTA 1E3HA 1KTVA 5ELWA 1N36I 4B3RI 4IZKB 4J3DA 2UUBE 1UDQA 4LFCE 4AIDA 3GMEA 4LFCI 3T1HI 5DRPA 1UDSA 4B3SI 2XEXA 4FW9A 4IZKA 4DV6I 2PNZA 2PNMA 1H7SA 4B3RE 5EKWA 2VF3A 3ZKDA 4WUCA 4LF6E 4IFDF 5DRQA 2BA0D 1XNQE 2Q2EB 2GO4A 4IFDE 4DV3E 4DR7E 1R6LA 1NHJA 4DU8A 4BA2A 3K1JA 3JAMQ 4HACA 1N32I 4JI0I 2V2VA 3V5RA 1RR9A 1XMOE 2UXDI 3P06A 2CZ9A 4IFDC 4N3OA 3ZZTA 1Z0BA 4YY3I 4I3HA 3J80Q 2AJ4A 2ZM6I 1XXEA 1IBKE 3T1YI 2CGEA 4X65I 4AIMA 4KHPI 3PYGA 1N32E 1HNZE 4OKGA 5L3JA 2UXCE 4IFDD 5EL9A 4Z7CA 1HNXE 4DV3I 3U1KA 4DR6E 1H7UA 4DPWA 3QT7A 4JI5E 4JG4A 2EFGA 4LF6I 2JEBA 3B78A 4X64I 2HHHE 4FWDA 4DR2E 3T1HE 1UDOA 1Z0VA 1EI1A 3M85G 1XNRI 4U3BA 4OO1A 2VQEI 1QZRA 2PNZB 5LL6R 5LMNE 3KRNA 4DUYI 1YH8A 2C38B 5FWKA 2JE6A 2UUCI 2GQ0A 1IBMI 1PIEA 3PYFA 1H73A 4OO1C 1KKHA 5DNXA 1N33I 4U3DA 2UUAE 1N0UA 4B3TI 4FWHA 3PS1A 3GCMA 2C38A 5JEAA 4LOMA 2Z5BA 5C0XD 1S4EA 4Z7YA 2GO3A 3GLLA 4R1FA 1DARA 4FW4A 5K36C 3PS3A 4JI8E 2E5LE 4JV5E 3U1YA 1Z5BA 2PO0B 4DPXA 5C0XC 2JEAB 3P3EA 2ZBKB 1HR0E 4DPUA 4MU4A 5JEAF 5C0XA 2PO2A 1Z0WA 2JT2A 3Q6MA 5JEAE 4LF4E 4IFDB 4BA1A 3OTOI 4IZJB 1UDNA 2WP8B 1HR0I 2F1DA 4DR1I 4P7AA 4YHHE 4JV5I 3H4LA 3ZZ0A 4DR2I 4JI2I 4DV1E 4DR5E 4JYAE 1IBLE 4AM3A 2AE8A 5IT9C 2IOPA 1ELOA 4DV0E 2R42A 1KVKA 4XTJA 2NN6F 4JI8I 2UUAI 1USUA 1NHIA 4IZJD 4URLA 3WU6A 3PYEA 4X65E 4GKJE 2ZITA 3CWVA 2BA0G 1IBKI 2IOQA 4JI2E 1H72C 3P3GA 2O1VA 5DRRA 1Z0TA 2UUCE 4AQYI 1ZM2A 4YHHI 1NHHA 2UXDE 4NBQA 3M7ND 1ZXMA 4MU1A 4LF7I 2F4VE 3WU3A 1VISA 4NXNI 4JI3I 3J81C 4LCGA 2C39A 4ED4A 1P42A 4MU0A 2OI2A 2VQEE 2BR2B 2UU9I 1Z0GA 1USVA 2NN6A 4B3SE 1KIJA 1R6MA 2UUBI 4LFAI 4M1KA 1E3PA 1N36E 3T1YE 1OYRA 4DR3E 5FLXQ 2NN6C 4YY3E 4BA1B 3JAMC 4K0KI 3L7ZA 2NPFA 2NN6B 1XMQE 2BA1D 4MYTA 1ZM3A 1I94I 4JI0E 3QT6A 4JI1E 2HFUA 1FJGE 4LCFA 2NN6E 4HYMA 1HK7A 2JEAA 2PO1A 2R3VA 1B63A 3LTOA 4MQYA 4OX9E 1XNQI 4FN5A 5C0XF 4LF9E 5K36A 2VESA 5K36D 1B62A 4PU9A 5C0XE 1XNRE 4LFAE 4DU7A 1RREA 2IERA 2LJPA 2V34A 4ZVIA 4LPFA 5K36B 2J7KA 4X64E 2PNLA 1ZM9A 1FKAE 5IT9Q 2HFSA 4DV6E 4JI5I 1MU5A 4OO1D 4QNJA 5IWAE 1ZXNA 1Z59A 1HNXI 1MX0A 2E1RA 2C37B 4UTGA 4MYUA 3M85D 4GKKE 4JI6I 3P76A 3B4TA 3HULA 1XMQI 4OO1B 1OJ4A 4USKA 1ZM4A 4DV4E 3ZKBA 4DR4E 4PRVA 4DR4I 3B8HA 4RPFA 2Z5CA 4DR3I 2V8PA 4JI1I 1XMOI 2VQFE 4OO1E 5DROA 4GKJI 1I94E 3LNUA 4JI6E 4OO1F 1RHYA 4ISAA 4IZJA 3B82A 2HK2A 5BR8E 2A2CA 4DV0I 1VI7A 3D4JA 2BR2A 3UHMA 2CVEA 4DPYA 3M7NG 2C37A 5K36E 4WUDA 2UXBE 1Y5YA 5K36F 1Y4UA 1J5EI 1FWLA 1NZ0A 4JI3E 4FW6A 5GMEA 3H1CA 4LF7E 1S16A 2NN6D 4DR1E 4LFBI 4LF9I 4DV7E 4NXME 1Z5AA 5JEAC 3M6AA 3K17A 4MDTA 3OTOE 1A6FA 5JEAB 2PO0A 4NXNE 4LF8E 1BKNA 2F4VI 4JI4E 4P52A 3J80C 3V2UC
chains in the Genus database with same CATH topology
3P76A 4U3DA 4J3DA 2JT2A 3PS1A 2O3ZA 1YHCA 3P3CA 2IERA 4LCGA 5DRPA 4FW6A 2IESA 1P42A 1XXEA 4IS9A 4OZEA 4U3BA 5DROA 2J65A 3PS2A 4LCHA 2GO3A 4OKGA 4ISAA 5DRQA 4LCFA 4FW4A 3P3GA 3PS3A 5DRRA 4MDTA 2GO4A 1YH8A 3U1YA 4FW3A 4FW7A 4MQYA 3NZKA 4FW5A 3P3EA 3UHMA 2VESA
chains in the Genus database with same CATH homology


 
#chains in the Genus database with same CATH superfamily
 3P76 A;  4U3D A;  4J3D A;  2JT2 A;  3PS1 A;  2O3Z A;  1YHC A;  3P3C A;  2IER A;  4LCG A;  5DRP A;  4FW6 A;  2IES A;  1P42 A;  1XXE A;  4IS9 A;  4OZE A;  4U3B A;  5DRO A;  2J65 A;  3PS2 A;  4LCH A;  2GO3 A;  4OKG A;  4ISA A;  5DRQ A;  4LCF A;  4FW4 A;  3P3G A;  3PS3 A;  5DRR A;  4MDT A;  2GO4 A;  1YH8 A;  3U1Y A;  4FW3 A;  4FW7 A;  4MQY A;  3NZK A;  4FW5 A;  3P3E A;  3UHM A;  2VES A; 
#chains in the Genus database with same CATH topology
 1Y6Z A;  3J7A M;  3HKM A;  2PO1 B;  2JE6 B;  1WUU A;  2BA1 G;  2GS8 A;  5LMU I;  1H74 A;  1OYP A;  2A2D A;  2V2Q A;  4DR5 I;  4DV5 I;  1D6T A;  4KQV A;  4DV2 I;  4EMD A;  3DD6 A;  1PVG A;  2UU9 E;  4X62 I;  2BM0 A;  1U2R A;  4B3M I;  4JYA I;  1HNW E;  4JI7 E;  2WP8 A;  4DV1 I;  4DUZ I;  1IBM E;  1N34 I;  5DNL A;  4LF5 I;  3Q6N A;  1YHC A;  1PKP A;  4PRX A;  2O1W A;  2DEI A;  1N34 E;  2HKJ A;  2WW4 A;  1XHK A;  2HK3 A;  4DXL A;  2J65 A;  4WUB A;  2Z5C B;  2X7I A;  4B3M E;  3WU5 A;  3PYD A;  3J7A G;  5GMD A;  1FNM A;  3K85 A;  1J5E E;  2O1T A;  4AQY E;  4QNK A;  4FW7 A;  5C0X B;  4DV5 E;  1K47 A;  3QT8 A;  2C39 B;  1Z5C A;  1Z0E A;  5BR8 I;  1MG7 A;  3P3C A;  2E5L I;  2UXC I;  4IS9 A;  1IBL I;  1N33 E;  5FLX C;  3ZZU A;  5LMU E;  4USM A;  2BV3 A;  4FW3 A;  3QT5 A;  2V2Z A;  4FW5 A;  5LMN I;  3PRY A;  1Z0C A;  1OYS A;  3CDI A;  3CDJ A;  1FI4 A;  1UEK A;  4MU3 A;  2O3Z A;  5JEA D;  4DR7 I;  3HJC A;  4LF4 I;  5IWA I;  2IES A;  3GON A;  4LF5 E;  2UXB I;  3LPS A;  4OZE A;  4DUZ E;  3F0N A;  4DV7 I;  4NXM I;  4HY1 A;  4JI7 I;  3PS2 A;  4LFB E;  4X62 E;  1HNW I;  3L7Z B;  2WNR B;  5A2Q Q;  4IFD A;  2X36 A;  3ZM7 A;  1N0V C;  4DV2 E;  3NZK A;  5A2Q C;  3Q1Q A;  2BM1 A;  4HXZ A;  1HNZ I;  2O1U A;  1FWK A;  2HHH I;  1Y60 A;  2CG9 A;  2VQF I;  2JEB B;  4OX9 I;  4GQU A;  4DR6 I;  1EA6 A;  4X66 I;  2Z5E A;  2WNR A;  2PO2 B;  3WU4 A;  4B3T E;  1FJG I;  4FWG A;  4KHP E;  4LCH A;  4HAC B;  4BA2 B;  4GKK I;  4UT4 A;  2DEJ A;  2Z5B B;  4JI4 I;  2ZM6 E;  4LF8 I;  1Y4S A;  4K0K E;  2HKE A;  4DUY E;  4X66 E;  4DV4 I;  3J81 Q;  4DPT A;  1E3H A;  1KTV A;  5ELW A;  1N36 I;  4B3R I;  4IZK B;  4J3D A;  2UUB E;  1UDQ A;  4LFC E;  4AID A;  3GME A;  4LFC I;  3T1H I;  5DRP A;  1UDS A;  4B3S I;  2XEX A;  4FW9 A;  4IZK A;  4DV6 I;  2PNZ A;  2PNM A;  1H7S A;  4B3R E;  5EKW A;  2VF3 A;  3ZKD A;  4WUC A;  4LF6 E;  4IFD F;  5DRQ A;  2BA0 D;  1XNQ E;  2Q2E B;  2GO4 A;  4IFD E;  4DV3 E;  4DR7 E;  1R6L A;  1NHJ A;  4DU8 A;  4BA2 A;  3K1J A;  3JAM Q;  4HAC A;  1N32 I;  4JI0 I;  2V2V A;  3V5R A;  1RR9 A;  1XMO E;  2UXD I;  3P06 A;  2CZ9 A;  4IFD C;  4N3O A;  3ZZT A;  1Z0B A;  4YY3 I;  4I3H A;  3J80 Q;  2AJ4 A;  2ZM6 I;  1XXE A;  1IBK E;  3T1Y I;  2CGE A;  4X65 I;  4AIM A;  4KHP I;  3PYG A;  1N32 E;  1HNZ E;  4OKG A;  5L3J A;  2UXC E;  4IFD D;  5EL9 A;  4Z7C A;  1HNX E;  4DV3 I;  3U1K A;  4DR6 E;  1H7U A;  4DPW A;  3QT7 A;  4JI5 E;  4JG4 A;  2EFG A;  4LF6 I;  2JEB A;  3B78 A;  4X64 I;  2HHH E;  4FWD A;  4DR2 E;  3T1H E;  1UDO A;  1Z0V A;  1EI1 A;  3M85 G;  1XNR I;  4U3B A;  4OO1 A;  2VQE I;  1QZR A;  2PNZ B;  5LL6 R;  5LMN E;  3KRN A;  4DUY I;  1YH8 A;  2C38 B;  5FWK A;  2JE6 A;  2UUC I;  2GQ0 A;  1IBM I;  1PIE A;  3PYF A;  1H73 A;  4OO1 C;  1KKH A;  5DNX A;  1N33 I;  4U3D A;  2UUA E;  1N0U A;  4B3T I;  4FWH A;  3PS1 A;  3GCM A;  2C38 A;  5JEA A;  4LOM A;  2Z5B A;  5C0X D;  1S4E A;  4Z7Y A;  2GO3 A;  3GLL A;  4R1F A;  1DAR A;  4FW4 A;  5K36 C;  3PS3 A;  4JI8 E;  2E5L E;  4JV5 E;  3U1Y A;  1Z5B A;  2PO0 B;  4DPX A;  5C0X C;  2JEA B;  3P3E A;  2ZBK B;  1HR0 E;  4DPU A;  4MU4 A;  5JEA F;  5C0X A;  2PO2 A;  1Z0W A;  2JT2 A;  3Q6M A;  5JEA E;  4LF4 E;  4IFD B;  4BA1 A;  3OTO I;  4IZJ B;  1UDN A;  2WP8 B;  1HR0 I;  2F1D A;  4DR1 I;  4P7A A;  4YHH E;  4JV5 I;  3H4L A;  3ZZ0 A;  4DR2 I;  4JI2 I;  4DV1 E;  4DR5 E;  4JYA E;  1IBL E;  4AM3 A;  2AE8 A;  5IT9 C;  2IOP A;  1ELO A;  4DV0 E;  2R42 A;  1KVK A;  4XTJ A;  2NN6 F;  4JI8 I;  2UUA I;  1USU A;  1NHI A;  4IZJ D;  4URL A;  3WU6 A;  3PYE A;  4X65 E;  4GKJ E;  2ZIT A;  3CWV A;  2BA0 G;  1IBK I;  2IOQ A;  4JI2 E;  1H72 C;  3P3G A;  2O1V A;  5DRR A;  1Z0T A;  2UUC E;  4AQY I;  1ZM2 A;  4YHH I;  1NHH A;  2UXD E;  4NBQ A;  3M7N D;  1ZXM A;  4MU1 A;  4LF7 I;  2F4V E;  3WU3 A;  1VIS A;  4NXN I;  4JI3 I;  3J81 C;  4LCG A;  2C39 A;  4ED4 A;  1P42 A;  4MU0 A;  2OI2 A;  2VQE E;  2BR2 B;  2UU9 I;  1Z0G A;  1USV A;  2NN6 A;  4B3S E;  1KIJ A;  1R6M A;  2UUB I;  4LFA I;  4M1K A;  1E3P A;  1N36 E;  3T1Y E;  1OYR A;  4DR3 E;  5FLX Q;  2NN6 C;  4YY3 E;  4BA1 B;  3JAM C;  4K0K I;  3L7Z A;  2NPF A;  2NN6 B;  1XMQ E;  2BA1 D;  4MYT A;  1ZM3 A;  1I94 I;  4JI0 E;  3QT6 A;  4JI1 E;  2HFU A;  1FJG E;  4LCF A;  2NN6 E;  4HYM A;  1HK7 A;  2JEA A;  2PO1 A;  2R3V A;  1B63 A;  3LTO A;  4MQY A;  4OX9 E;  1XNQ I;  4FN5 A;  5C0X F;  4LF9 E;  5K36 A;  2VES A;  5K36 D;  1B62 A;  4PU9 A;  5C0X E;  1XNR E;  4LFA E;  4DU7 A;  1RRE A;  2IER A;  2LJP A;  2V34 A;  4ZVI A;  4LPF A;  5K36 B;  2J7K A;  4X64 E;  2PNL A;  1ZM9 A;  1FKA E;  5IT9 Q;  2HFS A;  4DV6 E;  4JI5 I;  1MU5 A;  4OO1 D;  4QNJ A;  5IWA E;  1ZXN A;  1Z59 A;  1HNX I;  1MX0 A;  2E1R A;  2C37 B;  4UTG A;  4MYU A;  3M85 D;  4GKK E;  4JI6 I;  3P76 A;  3B4T A;  3HUL A;  1XMQ I;  4OO1 B;  1OJ4 A;  4USK A;  1ZM4 A;  4DV4 E;  3ZKB A;  4DR4 E;  4PRV A;  4DR4 I;  3B8H A;  4RPF A;  2Z5C A;  4DR3 I;  2V8P A;  4JI1 I;  1XMO I;  2VQF E;  4OO1 E;  5DRO A;  4GKJ I;  1I94 E;  3LNU A;  4JI6 E;  4OO1 F;  1RHY A;  4ISA A;  4IZJ A;  3B82 A;  2HK2 A;  5BR8 E;  2A2C A;  4DV0 I;  1VI7 A;  3D4J A;  2BR2 A;  3UHM A;  2CVE A;  4DPY A;  3M7N G;  2C37 A;  5K36 E;  4WUD A;  2UXB E;  1Y5Y A;  5K36 F;  1Y4U A;  1J5E I;  1FWL A;  1NZ0 A;  4JI3 E;  4FW6 A;  5GME A;  3H1C A;  4LF7 E;  1S16 A;  2NN6 D;  4DR1 E;  4LFB I;  4LF9 I;  4DV7 E;  4NXM E;  1Z5A A;  5JEA C;  3M6A A;  3K17 A;  4MDT A;  3OTO E;  1A6F A;  5JEA B;  2PO0 A;  4NXN E;  4LF8 E;  1BKN A;  2F4V I;  4JI4 E;  4P52 A;  3J80 C;  3V2U C; 
#chains in the Genus database with same CATH homology
 3P76 A;  4U3D A;  4J3D A;  2JT2 A;  3PS1 A;  2O3Z A;  1YHC A;  3P3C A;  2IER A;  4LCG A;  5DRP A;  4FW6 A;  2IES A;  1P42 A;  1XXE A;  4IS9 A;  4OZE A;  4U3B A;  5DRO A;  2J65 A;  3PS2 A;  4LCH A;  2GO3 A;  4OKG A;  4ISA A;  5DRQ A;  4LCF A;  4FW4 A;  3P3G A;  3PS3 A;  5DRR A;  4MDT A;  2GO4 A;  1YH8 A;  3U1Y A;  4FW3 A;  4FW7 A;  4MQY A;  3NZK A;  4FW5 A;  3P3E A;  3UHM A;  2VES A; 
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similar chains in the Genus database (?% sequence similarity)
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similar chains in the pdb database (?% sequence similarity)

 
#similar chains in the Genus database (?% sequence similarity)
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#similar chains, but unknotted
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#similar chains in the pdb database (?% sequence similarity)
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