1YHCA

Crystal structure of aquifex aeolicus lpxc deacetylase complexed with cacodylate
Total Genus 91
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The genus trace: a function that shows values of genus (vertical axis) for subchains spanned between the first residue, and all other residues (shown on horizontal axis). The number of the latter residue and the genus of a given subchain are shown interactively.

Total Genus
91
sequence length
266
structure length
266
Chain Sequence
GLEKTVKEKLSFEGVGIHTGEYSKLIIHPEKEGTGIRFFKNGVYIPARHEFVVHTNHSTDLGFKGQRIKTVEHILSVLHLLEITNVTIEVIGNEIPILDGSGWEFYEAIRKNILNQNREIDYFVVEEPIIVEDEGRLIKAEPSDTLEVTYEGEFKNFLGRQKFTFVEGNEEEIVLARTFAFDWEIEHIKKVGLGKGGSLKNTLVLGKDKVYNPEGLRYENEPVRHKVFDLIGDLYLLGSPVKGKFYSFRGGHSLNVKLVKELAKKQ
Loading diagram...

The genus matrix. At position (x,y) a genus value for a subchain spanned between x’th and y’th residue is shown. Values of the genus are represented by color, according to the scale given on the right.

Structure visualization

After clicking on a point (x,y) in the genus matrix above, a subchain from x to y is shown in color.

Chord Diagram
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molecule keywords UDP-3-O-[3-hydroxymyristoyl] N-acetylglucosamine deacetylase
publication title UDP-3-O-((R)-3-hydroxymyristoyl)-N-acetylglucosamine Deacetylase Functions through a General Acid-Base Catalyst Pair Mechanism
pubmed doi rcsb
source organism Aquifex aeolicus
molecule tags Hydrolase
total genus 91
structure length 266
sequence length 266
chains with identical sequence B
ec nomenclature ec 3.5.1.108: UDP-3-O-acyl-N-acetylglucosamine deacetylase.
pdb deposition date 2005-01-07

pfam database annotations

chain Pfam Accession Code Pfam Family Identifier Pfam Description
A PF03331 LpxC UDP-3-O-acyl N-acetylglycosamine deacetylase
Image from the rcsb pdb (www.rcsb.org)
cath code
ClassArchitectureTopologyHomologyDomain
3.30.230.20 Alpha Beta 2-Layer Sandwich Ribosomal Protein S5; domain 2 lpxc deacetylase, domain 1 1yhcA01
3.30.1700.10 Alpha Beta 2-Layer Sandwich lpxc deacetylase, domain 1 lpxc deacetylase, domain 2 1yhcA02
3UHMA 3PS1A 4FW5A 2GO3A 4FW7A 1XXEA 4FW3A 3P76A 1YH8A 3U1YA 4U3DA 2IERA 2J65A 2JT2A 1YHCA 5DRQA 4FW4A 2IESA 4IS9A 4MQYA 4MDTA 4FW6A 1P42A 4J3DA 4OKGA 4LCGA 3NZKA 3PS3A 2VESA 3P3CA 5DROA 4LCHA 4ISAA 2GO4A 2O3ZA 3P3EA 4U3BA 4LCFA 5DRPA 4OZEA 5DRRA 3P3GA 3PS2A
chains in the Genus database with same CATH superfamily
5EKWA 2JEBB 1N0UA 1OYPA 3B82A 1UDQA 4DV6E 4MU1A 1XMOE 4DUZE 4X62E 2PO0B 4Z7YA 4JI3I 4OO1D 4RPFA 4HYMA 4IS9A 5IT9Q 4J3DA 3D4JA 5IT9C 1R6LA 4BA2A 2Z5BB 2UUCI 4DR3E 4MYTA 5GMEA 2O3ZA 4AIMA 4DR6I 4DR7E 5K36A 4NBQA 4DPUA 2UXBI 3QT6A 3QT7A 1ZM3A 3ZKBA 4BA1B 3QT5A 3QT8A 5LMUE 2HFUA 3U1YA 2HKJA 4DXLA 4USMA 5JEAF 4X66I 1XNQI 4JI4I 1N36I 1IBKI 4DUYI 4DPXA 4M1KA 4JG4A 5IWAE 4DV2I 2PNZA 4B3SI 1E3PA 3PYDA 4LF9I 1H74A 2PO2A 4DV5E 2CVEA 5BR8E 4JI6I 5JEAB 2HK3A 2NN6F 3K85A 3OTOI 2PO1B 4B3RE 1Z0TA 3L7ZB 3B78A 2F4VI 1Z0VA 4DV6I 3M7NG 2WNRB 4EMDA 2BA1G 4DV0E 1OJ4A 4AM3A 1HR0E 5DNLA 3WU5A 2C37B 4WUBA 1N32E 1FI4A 5JEAD 3WU6A 2NN6B 3P3CA 4DR3I 1E3HA 4HY1A 4DR2E 2UXDE 2UXCI 4DR1I 3U1KA 5A2QC 5DNXA 4DV7E 1EA6A 2BR2B 4GKKI 3V2UC 2V34A 4IFDF 2NN6D 4JI1I 2ZM6I 3J7AG 4DR4E 5DRQA 1XHKA 1Y4SA 4MQYA 4JI7E 3LTOA 3PRYA 4YHHE 3PS3A 4DR5E 2O1TA 4NXME 4LPFA 1H72C 1UDSA 1UDOA 4LCHA 2C39B 4FWHA 2VQFI 4IFDB 1IBMI 3Q6NA 4P52A 5JEAE 1J5EE 5DRPA 4MU4A 3PS2A 4MYUA 2IOPA 4B3RI 2E5LE 4JYAE 2GO3A 4QNKA 4QNJA 3P76A 3T1HI 4URLA 4LF8E 3PYEA 2C39A 4IFDD 4DV4E 4GKJE 4IFDA 4OO1C 2NN6E 2BV3A 4IZJA 1USVA 1FJGI 4LFAE 1Z5BA 5K36F 5DROA 1HNZE 3DD6A 4KHPI 4DR2I 3P3EA 3T1YE 5C0XC 2R42A 4BA2B 2BA0D 4DPTA 3LNUA 1Z0CA 4LF7I 5K36B 1Z0GA 4X64I 4LF5E 2OI2A 4FW4A 2WW4A 4LCGA 1N33I 2HHHI 2VQEE 5EL9A 4DV1I 4FWDA 4IFDE 2UUBI 1KKHA 2E1RA 3CDIA 5K36D 2PNZB 2V2VA 3P3GA 3ZZTA 4HACA 4I3HA 3PYGA 2PO2B 4HXZA 4P7AA 1FWKA 1XXEA 4LF4I 2R3VA 4MU0A 3HJCA 2PNMA 4AQYI 4R1FA 3V5RA 4X65I 2DEIA 4DV3I 4DV4I 1KTVA 1ZM2A 2JEAA 3J80Q 3J80C 1Z0BA 2V8PA 4ISAA 2A2CA 3F0NA 1NZ0A 1Z0EA 1HNZI 3GCMA 1Z5AA 2UU9E 4NXNE 4B3TE 1RREA 4B3ME 3CWVA 1Y4UA 3WU4A 4Z7CA 4YY3E 3JAMQ 4UT4A 5K36E 3K17A 3JAMC 5LL6R 1XMOI 1MX0A 4DUZI 4X62I 4JI5E 4JV5E 2JT2A 1HNWE 4FN5A 4UTGA 1YHCA 2EFGA 4LF5I 1QZRA 4LFBE 1XNRE 2GQ0A 1RHYA 3CDJA 4JI8E 1D6TA 4LF6E 3HKMA 2A2DA 5FWKA 2VQEI 1ZXMA 1R6MA 2DEJA 5L3JA 2WP8A 1Z59A 1N34E 5A2QQ 4DR7I 1FWLA 2O1VA 4LFCE 4JI2E 3KRNA 1OYSA 5LMUI 2JE6A 1MU5A 1I94E 4U3DA 2UUAE 1B63A 4FW9A 3ZZ0A 1Z0WA 4JI0E 1IBLE 2Z5CB 3K1JA 5IWAI 1ZXNA 4USKA 1NHHA 5FLXC 4KQVA 1HNXE 2AJ4A 1XMQE 3M85D 4DV5I 5BR8I 4NXNI 4K0KE 4IZJB 2V2ZA 1KVKA 3UHMA 2CG9A 4PRXA 2Z5CA 2Q2EB 4JI3E 2X7IA 1UEKA 1HNWI 2V2QA 4DV0I 4WUCA 3H1CA 1HR0I 1N32I 4IZJD 2UUCE 1H7SA 3P06A 3J81C 4DR6E 4DPWA 2UXBE 2UXDI 5DRRA 5K36C 1DARA 2BM1A 4OO1F 4DV7I 4IZKB 1H73A 4FW3A 3Q6MA 1XNQE 2CZ9A 2IERA 4JI4E 4AIDA 1N36E 1IBKE 4DUYE 4HACB 3PYFA 4WUDA 1USUA 2GS8A 4DV2E 4DR4I 4B3SE 5LMNE 5C0XF 1IBLI 2PNLA 4LF9E 1WUUA 4JI7I 1VISA 4YHHI 4DR5I 4NXMI 2GO4A 3ZZUA 4IZKA 1RR9A 1XMQI 1J5EI 3OTOE 2JEAB 2C38A 4DU7A 3GONA 3H4LA 2F4VE 2AE8A 5C0XB 4PU9A 2E5LI 3M7ND 4JYAI 1VI7A 2BA1D 1ZM9A 4OO1A 4LF8I 1KIJA 4FW6A 2X36A 4FWGA 4GKJI 4OKGA 1Y6ZA 2HK2A 2JEBA 5JEAC 4LFAI 5C0XD 2UXCE 5C0XA 4LOMA 4DR1E 4OX9E 2PO0A 3T1YI 1B62A 3Q1QA 1Y60A 2BA0G 4GKKE 1S16A 2WP8B 4JI1E 2Z5BA 2BM0A 2ZM6E 4ED4A 2NN6C 1PKPA 5LMNI 3HULA 4MDTA 3NZKA 1BKNA 2JE6B 2VESA 4OO1E 4BA1A 2VQFE 1IBME 1Y5YA 4U3BA 2ZITA 4OZEA 5C0XE 1PVGA 4XTJA 4FW7A 3T1HE 1YH8A 1OYRA 1PIEA 2J65A 4DU8A 3ZM7A 3GLLA 3WU3A 1FJGE 5JEAA 1Z5CA 1P42A 4ZVIA 2VF3A 1N0VC 4DPYA 2PO1A 4IFDC 5FLXQ 3L7ZA 5ELWA 4KHPE 3LPSA 2WNRA 4OX9I 1ELOA 4N3OA 2UU9I 4B3TI 4B3MI 2C37A 2IOQA 1FNMA 3PS1A 1FKAE 4FW5A 2CGEA 1S4EA 4YY3I 4LF7E 2J7KA 1NHIA 2NN6A 3B8HA 4X64E 2HFSA 1U2RA 4JI5I 4JV5I 2NPFA 2O1WA 2O1UA 2IESA 1K47A 4LFBI 1N33E 1XNRI 2HHHE 4JI8I 4DV1E 3J7AM 4LF6I 2XEXA 2ZBKB 3J81Q 2UUBE 2BR2A 3GMEA 3B4TA 4PRVA 3ZKDA 4LCFA 1A6FA 1N34I 2Z5EA 1EI1A 4LF4E 4LFCI 4JI2I 4GQUA 4AQYE 1H7UA 4X66E 1HK7A 1UDNA 1I94I 2UUAI 2LJPA 4X65E 2HKEA 2C38B 4DV3E 4JI0I 1MG7A 2F1DA 5GMDA 1ZM4A 4MU3A 4OO1B 3M6AA 3M85G 1HNXI 4JI6E 1NHJA 4K0KI
chains in the Genus database with same CATH topology
3UHMA 3PS1A 4FW5A 2GO3A 4FW7A 1XXEA 4FW3A 3P76A 1YH8A 3U1YA 4U3DA 2IERA 2J65A 2JT2A 1YHCA 5DRQA 4FW4A 2IESA 4IS9A 4MQYA 4MDTA 4FW6A 1P42A 4J3DA 4OKGA 4LCGA 3NZKA 3PS3A 2VESA 3P3CA 5DROA 4LCHA 4ISAA 2GO4A 2O3ZA 3P3EA 4U3BA 4LCFA 5DRPA 4OZEA 5DRRA 3P3GA 3PS2A
chains in the Genus database with same CATH homology


 
#chains in the Genus database with same CATH superfamily
 3UHM A;  3PS1 A;  4FW5 A;  2GO3 A;  4FW7 A;  1XXE A;  4FW3 A;  3P76 A;  1YH8 A;  3U1Y A;  4U3D A;  2IER A;  2J65 A;  2JT2 A;  1YHC A;  5DRQ A;  4FW4 A;  2IES A;  4IS9 A;  4MQY A;  4MDT A;  4FW6 A;  1P42 A;  4J3D A;  4OKG A;  4LCG A;  3NZK A;  3PS3 A;  2VES A;  3P3C A;  5DRO A;  4LCH A;  4ISA A;  2GO4 A;  2O3Z A;  3P3E A;  4U3B A;  4LCF A;  5DRP A;  4OZE A;  5DRR A;  3P3G A;  3PS2 A; 
#chains in the Genus database with same CATH topology
 5EKW A;  2JEB B;  1N0U A;  1OYP A;  3B82 A;  1UDQ A;  4DV6 E;  4MU1 A;  1XMO E;  4DUZ E;  4X62 E;  2PO0 B;  4Z7Y A;  4JI3 I;  4OO1 D;  4RPF A;  4HYM A;  4IS9 A;  5IT9 Q;  4J3D A;  3D4J A;  5IT9 C;  1R6L A;  4BA2 A;  2Z5B B;  2UUC I;  4DR3 E;  4MYT A;  5GME A;  2O3Z A;  4AIM A;  4DR6 I;  4DR7 E;  5K36 A;  4NBQ A;  4DPU A;  2UXB I;  3QT6 A;  3QT7 A;  1ZM3 A;  3ZKB A;  4BA1 B;  3QT5 A;  3QT8 A;  5LMU E;  2HFU A;  3U1Y A;  2HKJ A;  4DXL A;  4USM A;  5JEA F;  4X66 I;  1XNQ I;  4JI4 I;  1N36 I;  1IBK I;  4DUY I;  4DPX A;  4M1K A;  4JG4 A;  5IWA E;  4DV2 I;  2PNZ A;  4B3S I;  1E3P A;  3PYD A;  4LF9 I;  1H74 A;  2PO2 A;  4DV5 E;  2CVE A;  5BR8 E;  4JI6 I;  5JEA B;  2HK3 A;  2NN6 F;  3K85 A;  3OTO I;  2PO1 B;  4B3R E;  1Z0T A;  3L7Z B;  3B78 A;  2F4V I;  1Z0V A;  4DV6 I;  3M7N G;  2WNR B;  4EMD A;  2BA1 G;  4DV0 E;  1OJ4 A;  4AM3 A;  1HR0 E;  5DNL A;  3WU5 A;  2C37 B;  4WUB A;  1N32 E;  1FI4 A;  5JEA D;  3WU6 A;  2NN6 B;  3P3C A;  4DR3 I;  1E3H A;  4HY1 A;  4DR2 E;  2UXD E;  2UXC I;  4DR1 I;  3U1K A;  5A2Q C;  5DNX A;  4DV7 E;  1EA6 A;  2BR2 B;  4GKK I;  3V2U C;  2V34 A;  4IFD F;  2NN6 D;  4JI1 I;  2ZM6 I;  3J7A G;  4DR4 E;  5DRQ A;  1XHK A;  1Y4S A;  4MQY A;  4JI7 E;  3LTO A;  3PRY A;  4YHH E;  3PS3 A;  4DR5 E;  2O1T A;  4NXM E;  4LPF A;  1H72 C;  1UDS A;  1UDO A;  4LCH A;  2C39 B;  4FWH A;  2VQF I;  4IFD B;  1IBM I;  3Q6N A;  4P52 A;  5JEA E;  1J5E E;  5DRP A;  4MU4 A;  3PS2 A;  4MYU A;  2IOP A;  4B3R I;  2E5L E;  4JYA E;  2GO3 A;  4QNK A;  4QNJ A;  3P76 A;  3T1H I;  4URL A;  4LF8 E;  3PYE A;  2C39 A;  4IFD D;  4DV4 E;  4GKJ E;  4IFD A;  4OO1 C;  2NN6 E;  2BV3 A;  4IZJ A;  1USV A;  1FJG I;  4LFA E;  1Z5B A;  5K36 F;  5DRO A;  1HNZ E;  3DD6 A;  4KHP I;  4DR2 I;  3P3E A;  3T1Y E;  5C0X C;  2R42 A;  4BA2 B;  2BA0 D;  4DPT A;  3LNU A;  1Z0C A;  4LF7 I;  5K36 B;  1Z0G A;  4X64 I;  4LF5 E;  2OI2 A;  4FW4 A;  2WW4 A;  4LCG A;  1N33 I;  2HHH I;  2VQE E;  5EL9 A;  4DV1 I;  4FWD A;  4IFD E;  2UUB I;  1KKH A;  2E1R A;  3CDI A;  5K36 D;  2PNZ B;  2V2V A;  3P3G A;  3ZZT A;  4HAC A;  4I3H A;  3PYG A;  2PO2 B;  4HXZ A;  4P7A A;  1FWK A;  1XXE A;  4LF4 I;  2R3V A;  4MU0 A;  3HJC A;  2PNM A;  4AQY I;  4R1F A;  3V5R A;  4X65 I;  2DEI A;  4DV3 I;  4DV4 I;  1KTV A;  1ZM2 A;  2JEA A;  3J80 Q;  3J80 C;  1Z0B A;  2V8P A;  4ISA A;  2A2C A;  3F0N A;  1NZ0 A;  1Z0E A;  1HNZ I;  3GCM A;  1Z5A A;  2UU9 E;  4NXN E;  4B3T E;  1RRE A;  4B3M E;  3CWV A;  1Y4U A;  3WU4 A;  4Z7C A;  4YY3 E;  3JAM Q;  4UT4 A;  5K36 E;  3K17 A;  3JAM C;  5LL6 R;  1XMO I;  1MX0 A;  4DUZ I;  4X62 I;  4JI5 E;  4JV5 E;  2JT2 A;  1HNW E;  4FN5 A;  4UTG A;  1YHC A;  2EFG A;  4LF5 I;  1QZR A;  4LFB E;  1XNR E;  2GQ0 A;  1RHY A;  3CDJ A;  4JI8 E;  1D6T A;  4LF6 E;  3HKM A;  2A2D A;  5FWK A;  2VQE I;  1ZXM A;  1R6M A;  2DEJ A;  5L3J A;  2WP8 A;  1Z59 A;  1N34 E;  5A2Q Q;  4DR7 I;  1FWL A;  2O1V A;  4LFC E;  4JI2 E;  3KRN A;  1OYS A;  5LMU I;  2JE6 A;  1MU5 A;  1I94 E;  4U3D A;  2UUA E;  1B63 A;  4FW9 A;  3ZZ0 A;  1Z0W A;  4JI0 E;  1IBL E;  2Z5C B;  3K1J A;  5IWA I;  1ZXN A;  4USK A;  1NHH A;  5FLX C;  4KQV A;  1HNX E;  2AJ4 A;  1XMQ E;  3M85 D;  4DV5 I;  5BR8 I;  4NXN I;  4K0K E;  4IZJ B;  2V2Z A;  1KVK A;  3UHM A;  2CG9 A;  4PRX A;  2Z5C A;  2Q2E B;  4JI3 E;  2X7I A;  1UEK A;  1HNW I;  2V2Q A;  4DV0 I;  4WUC A;  3H1C A;  1HR0 I;  1N32 I;  4IZJ D;  2UUC E;  1H7S A;  3P06 A;  3J81 C;  4DR6 E;  4DPW A;  2UXB E;  2UXD I;  5DRR A;  5K36 C;  1DAR A;  2BM1 A;  4OO1 F;  4DV7 I;  4IZK B;  1H73 A;  4FW3 A;  3Q6M A;  1XNQ E;  2CZ9 A;  2IER A;  4JI4 E;  4AID A;  1N36 E;  1IBK E;  4DUY E;  4HAC B;  3PYF A;  4WUD A;  1USU A;  2GS8 A;  4DV2 E;  4DR4 I;  4B3S E;  5LMN E;  5C0X F;  1IBL I;  2PNL A;  4LF9 E;  1WUU A;  4JI7 I;  1VIS A;  4YHH I;  4DR5 I;  4NXM I;  2GO4 A;  3ZZU A;  4IZK A;  1RR9 A;  1XMQ I;  1J5E I;  3OTO E;  2JEA B;  2C38 A;  4DU7 A;  3GON A;  3H4L A;  2F4V E;  2AE8 A;  5C0X B;  4PU9 A;  2E5L I;  3M7N D;  4JYA I;  1VI7 A;  2BA1 D;  1ZM9 A;  4OO1 A;  4LF8 I;  1KIJ A;  4FW6 A;  2X36 A;  4FWG A;  4GKJ I;  4OKG A;  1Y6Z A;  2HK2 A;  2JEB A;  5JEA C;  4LFA I;  5C0X D;  2UXC E;  5C0X A;  4LOM A;  4DR1 E;  4OX9 E;  2PO0 A;  3T1Y I;  1B62 A;  3Q1Q A;  1Y60 A;  2BA0 G;  4GKK E;  1S16 A;  2WP8 B;  4JI1 E;  2Z5B A;  2BM0 A;  2ZM6 E;  4ED4 A;  2NN6 C;  1PKP A;  5LMN I;  3HUL A;  4MDT A;  3NZK A;  1BKN A;  2JE6 B;  2VES A;  4OO1 E;  4BA1 A;  2VQF E;  1IBM E;  1Y5Y A;  4U3B A;  2ZIT A;  4OZE A;  5C0X E;  1PVG A;  4XTJ A;  4FW7 A;  3T1H E;  1YH8 A;  1OYR A;  1PIE A;  2J65 A;  4DU8 A;  3ZM7 A;  3GLL A;  3WU3 A;  1FJG E;  5JEA A;  1Z5C A;  1P42 A;  4ZVI A;  2VF3 A;  1N0V C;  4DPY A;  2PO1 A;  4IFD C;  5FLX Q;  3L7Z A;  5ELW A;  4KHP E;  3LPS A;  2WNR A;  4OX9 I;  1ELO A;  4N3O A;  2UU9 I;  4B3T I;  4B3M I;  2C37 A;  2IOQ A;  1FNM A;  3PS1 A;  1FKA E;  4FW5 A;  2CGE A;  1S4E A;  4YY3 I;  4LF7 E;  2J7K A;  1NHI A;  2NN6 A;  3B8H A;  4X64 E;  2HFS A;  1U2R A;  4JI5 I;  4JV5 I;  2NPF A;  2O1W A;  2O1U A;  2IES A;  1K47 A;  4LFB I;  1N33 E;  1XNR I;  2HHH E;  4JI8 I;  4DV1 E;  3J7A M;  4LF6 I;  2XEX A;  2ZBK B;  3J81 Q;  2UUB E;  2BR2 A;  3GME A;  3B4T A;  4PRV A;  3ZKD A;  4LCF A;  1A6F A;  1N34 I;  2Z5E A;  1EI1 A;  4LF4 E;  4LFC I;  4JI2 I;  4GQU A;  4AQY E;  1H7U A;  4X66 E;  1HK7 A;  1UDN A;  1I94 I;  2UUA I;  2LJP A;  4X65 E;  2HKE A;  2C38 B;  4DV3 E;  4JI0 I;  1MG7 A;  2F1D A;  5GMD A;  1ZM4 A;  4MU3 A;  4OO1 B;  3M6A A;  3M85 G;  1HNX I;  4JI6 E;  1NHJ A;  4K0K I; 
#chains in the Genus database with same CATH homology
 3UHM A;  3PS1 A;  4FW5 A;  2GO3 A;  4FW7 A;  1XXE A;  4FW3 A;  3P76 A;  1YH8 A;  3U1Y A;  4U3D A;  2IER A;  2J65 A;  2JT2 A;  1YHC A;  5DRQ A;  4FW4 A;  2IES A;  4IS9 A;  4MQY A;  4MDT A;  4FW6 A;  1P42 A;  4J3D A;  4OKG A;  4LCG A;  3NZK A;  3PS3 A;  2VES A;  3P3C A;  5DRO A;  4LCH A;  4ISA A;  2GO4 A;  2O3Z A;  3P3E A;  4U3B A;  4LCF A;  5DRP A;  4OZE A;  5DRR A;  3P3G A;  3PS2 A; 
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similar chains in the Genus database (?% sequence similarity)
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similar chains in the pdb database (?% sequence similarity)

 
#similar chains in the Genus database (?% sequence similarity)
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#similar chains, but unknotted
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#similar chains in the pdb database (?% sequence similarity)
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