1YHCA

Crystal structure of aquifex aeolicus lpxc deacetylase complexed with cacodylate
Total Genus 91
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The genus trace: a function that shows values of genus (vertical axis) for subchains spanned between the first residue, and all other residues (shown on horizontal axis). The number of the latter residue and the genus of a given subchain are shown interactively.

Total Genus
91
sequence length
266
structure length
266
Chain Sequence
GLEKTVKEKLSFEGVGIHTGEYSKLIIHPEKEGTGIRFFKNGVYIPARHEFVVHTNHSTDLGFKGQRIKTVEHILSVLHLLEITNVTIEVIGNEIPILDGSGWEFYEAIRKNILNQNREIDYFVVEEPIIVEDEGRLIKAEPSDTLEVTYEGEFKNFLGRQKFTFVEGNEEEIVLARTFAFDWEIEHIKKVGLGKGGSLKNTLVLGKDKVYNPEGLRYENEPVRHKVFDLIGDLYLLGSPVKGKFYSFRGGHSLNVKLVKELAKKQ
Loading diagram...

The genus matrix. At position (x,y) a genus value for a subchain spanned between x’th and y’th residue is shown. Values of the genus are represented by color, according to the scale given on the right.

Structure visualization

After clicking on a point (x,y) in the genus matrix above, a subchain from x to y is shown in color.

Chord Diagram
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molecule keywords UDP-3-O-[3-hydroxymyristoyl] N-acetylglucosamine deacetylase
publication title UDP-3-O-((R)-3-hydroxymyristoyl)-N-acetylglucosamine Deacetylase Functions through a General Acid-Base Catalyst Pair Mechanism
pubmed doi rcsb
source organism Aquifex aeolicus
molecule tags Hydrolase
total genus 91
structure length 266
sequence length 266
chains with identical sequence B
ec nomenclature ec 3.5.1.108: UDP-3-O-acyl-N-acetylglucosamine deacetylase.
pdb deposition date 2005-01-07

pfam database annotations

chain Pfam Accession Code Pfam Family Identifier Pfam Description
A PF03331 LpxC UDP-3-O-acyl N-acetylglycosamine deacetylase
Image from the rcsb pdb (www.rcsb.org)
cath code
ClassArchitectureTopologyHomologyDomain
3.30.230.20 Alpha Beta 2-Layer Sandwich Ribosomal Protein S5; domain 2 lpxc deacetylase, domain 1 1yhcA01
3.30.1700.10 Alpha Beta 2-Layer Sandwich lpxc deacetylase, domain 1 lpxc deacetylase, domain 2 1yhcA02
3P76A 4LCFA 2J65A 4IS9A 4LCGA 4FW5A 2GO3A 1P42A 3PS2A 3P3CA 4FW7A 5DROA 2O3ZA 4U3BA 3NZKA 1YH8A 3U1YA 2IERA 4MDTA 3P3GA 5DRRA 4J3DA 1XXEA 4FW3A 4ISAA 4MQYA 3PS3A 4FW4A 5DRPA 4OKGA 5DRQA 4LCHA 3P3EA 4U3DA 4FW6A 3PS1A 2IESA 2VESA 2GO4A 3UHMA 4OZEA 2JT2A 1YHCA
chains in the Genus database with same CATH superfamily
5JEAE 1OYPA 4JI7I 1KIJA 4FW5A 3T1HE 5A2QC 2UUCE 3K17A 2F4VI 1R6MA 2C38B 4DV4E 2UXCE 4IFDA 4HACB 1YH8A 1UDSA 2NN6E 2ZM6I 1N36I 2UU9I 1XMOI 2JEAB 1S4EA 5L3JA 4KHPI 3M85D 4QNJA 4LF5E 4DR6I 3F0NA 4WUCA 4JI5E 2PO1A 1ZM9A 2V8PA 2VESA 2C37B 4X66I 2OI2A 2IOQA 5IWAI 1J5EE 2C39B 1H7SA 3ZKDA 2Z5EA 2HHHI 2BA0G 4JI0E 2XEXA 2PO2A 2V34A 2WW4A 4DV1I 4IZJA 4ED4A 2X36A 3Q6MA 4JG4A 4RPFA 4NXME 4P52A 4B3TI 5K36A 2WNRB 4DV6I 2UXDE 1USUA 2AJ4A 2JT2A 1MU5A 4DR2I 5C0XA 1NHIA 1H73A 5DROA 1HNXI 4PRXA 4DUYE 3JAMC 4WUDA 1IBLI 4DV2E 3U1YA 4UT4A 5DNXA 4IFDB 4DV3I 1IBKE 4UTGA 5FLXQ 1N34I 2E5LE 4MU3A 4DR3E 1Z0WA 4I3HA 1HNWI 2Z5CB 4JV5I 2BM1A 3OTOI 4IFDC 4B3SI 4LF4E 2O1VA 2PO1B 1XNRE 2NN6A 1UDQA 1XHKA 3U1KA 4EMDA 4OO1F 2GS8A 1FKAE 4JI4I 3HULA 1VISA 2CG9A 4DV0I 3J81C 2O1TA 4JYAI 1HNZE 2PO2B 2AE8A 4FW9A 4LF9E 4X64E 5LMNI 1PVGA 4M1KA 4MYTA 1IBMI 1ZM4A 4IZJB 4JI6I 4OX9I 3HKMA 3D4JA 4YY3I 1BKNA 1ELOA 4ZVIA 5K36B 5JEAF 5EL9A 2BA0D 4JI7E 4PU9A 4NBQA 2WP8A 3QT8A 3KRNA 2PNLA 5C0XB 2F4VE 1XNQI 3ZZTA 5LMUI 1N32I 3J80Q 2CVEA 4JI1I 4GKJI 2ZM6E 3DD6A 1N36E 2UU9E 1XMOE 4AIMA 2E1RA 4KHPE 3H1CA 4DR6E 2PO0A 2CGEA 1KTVA 4DPYA 4X66E 2UXBI 2VQFI 3B8HA 4IFDD 2IESA 5IWAE 3GONA 3B4TA 4DV1E 4JI3I 2PNMA 2NPFA 4LF7I 4DV7I 2Z5BB 3K85A 4AIDA 4URLA 4DPWA 5DRRA 4B3TE 4B3MI 4DV6E 2DEIA 4GKKI 2R3VA 5FWKA 3Q6NA 5LL6R 2O1WA 1USVA 4DR2E 4LFAI 1NHJA 2WP8B 2HKJA 3PS2A 1IBLE 4YHHI 2BR2A 3P06A 2UUAE 4DV3E 1I94I 4DPUA 2HK3A 1N34E 1FWLA 1HNWE 5C0XD 4JV5E 4OO1A 1ZM2A 2PO0B 4DR5I 1FNMA 4FW6A 4JI4E 4GQUA 4N3OA 4MYUA 2PNZA 1Z0CA 2UUBI 3P3CA 2ZBKB 1Z0VA 4LFCE 4DR7I 5LMNE 4DR4E 1IBME 4LF8I 2GQ0A 5FLXC 1Y60A 1XMQE 1KKHA 2VF3A 4AQYI 4JI6E 4OX9E 1Z5CA 4YY3E 3CDIA 1MG7A 5JEAA 1Y6ZA 4BA1A 2X7IA 2V2ZA 2JE6A 1N0VC 4OZEA 1XNQE 1ZM3A 1Z5BA 5LMUE 1N32E 3ZZUA 1J5EI 3M7NG 4JI1E 4MU4A 4DPXA 4GKJE 4DV5I 1HR0E 1WUUA 1Z0TA 1FJGI 5IT9Q 3WU6A 5BR8I 2HFSA 1H72C 3T1YI 4B3RI 1KVKA 3L7ZA 4OO1B 3WU5A 2VQFE 2EFGA 4X65I 2F1DA 4U3DA 2O1UA 1R6LA 4IFDE 3J7AG 4HYMA 4FWDA 2CZ9A 1H7UA 4JI3E 1MX0A 2PNZB 3J80C 3GLLA 4LF7E 3PYDA 5K36C 4DUZI 4DV7E 1FWKA 1PIEA 2O3ZA 4B3ME 4R1FA 1B63A 5C0XC 1HK7A 5JEAB 4GKKE 2V2VA 4BA1B 3WU3A 2V2QA 4JI2I 2C37A 4LFAE 2JE6B 4NXNI 4LF6I 2NN6B 2HHHE 1D6TA 2A2DA 2R42A 4WUBA 4YHHE 1P42A 2JEBA 2BV3A 2J7KA 1Z59A 1I94E 4DR1I 1Z0BA 2NN6C 1HNZI 1K47A 3P3GA 3GCMA 4IZKA 2VQEI 4LF9I 4X64I 4LOMA 2Q2EB 3PYFA 1N33I 3L7ZB 2BA1G 4MQYA 4XTJA 1E3PA 3CWVA 3M7ND 4DU8A 4BA2A 4IZJD 4K0KI 3H4LA 2UUBE 4LFBI 1YHCA 4MU1A 4OO1D 3LTOA 4LCFA 4DR7E 4IS9A 4HXZA 1HNXE 4JI8I 4LF8E 1Y4UA 5K36D 4DPTA 3ZM7A 4AQYE 3V5RA 5A2QQ 3GMEA 4X62I 4MU0A 3ZKBA 3J7AM 1Z0GA 3PYEA 2Z5CA 1Y5YA 3OTOE 4B3SE 4FW4A 1ZXMA 3ZZ0A 1OJ4A 2IOPA 2GO4A 1Z0EA 3LPSA 4AM3A 4DV0E 3LNUA 5JEAD 4DV5E 4JYAE 3B82A 4FW7A 1FJGE 2JEBB 5BR8E 3T1YE 4PRVA 4Z7YA 5ELWA 3T1HI 4B3RE 2UUCI 5GMEA 1RREA 2IERA 2NN6D 4MDTA 4DV4I 1OYRA 4IZKB 1UDNA 2UXCI 1UEKA 1XXEA 4X65E 4ISAA 3QT5A 4IFDF 1NZ0A 4LF5I 2LJPA 4JI5I 1N0UA 4HY1A 4BA2B 4DUZE 1EA6A 3P76A 3WU4A 2DEJA 2J65A 4LCGA 4FWGA 5IT9C 1ZXNA 1DARA 1H74A 4JI0I 2BR2B 3M6AA 2UUAI 4USMA 4USKA 1FI4A 1OYSA 3QT6A 2BA1D 3JAMQ 3PRYA 1B62A 4J3DA 4JI2E 4LPFA 5DRPA 2UXBE 4NXMI 4LCHA 4NXNE 3PYGA 4LF6E 3PS1A 3M85G 1Y4SA 2UXDI 4OO1C 5K36F 4DR1E 4Z7CA 2C38A 2A2CA 4KQVA 1Z5AA 2Z5BA 2ZITA 4QNKA 5EKWA 4LFCI 3J81Q 2VQEE 1UDOA 4HACA 4DXLA 4DR4I 4U3BA 3NZKA 5C0XF 4DUYI 1RHYA 1XMQI 3Q1QA 1N33E 4DV2I 3QT7A 2BM0A 2JEAA 1E3HA 1IBKI 3PS3A 1PKPA 1S16A 2E5LI 4OKGA 4DR3I 4K0KE 3B78A 4LFBE 4DU7A 4P7AA 3UHMA 4LF4I 5JEAC 2C39A 3CDJA 1XNRI 3K1JA 4JI8E 4FWHA 4OO1E 1U2RA 5DNLA 2GO3A 2HFUA 2HKEA 2NN6F 2HK2A 5K36E 1HR0I 4X62E 1QZRA 1NHHA 1EI1A 3HJCA 4FW3A 1VI7A 5C0XE 2WNRA 4DR5E 5DRQA 3P3EA 5GMDA 1RR9A 1A6FA 4FN5A 3V2UC
chains in the Genus database with same CATH topology
3P76A 4LCFA 2J65A 4IS9A 4LCGA 4FW5A 2GO3A 1P42A 3PS2A 3P3CA 4FW7A 5DROA 2O3ZA 4U3BA 3NZKA 1YH8A 3U1YA 2IERA 4MDTA 3P3GA 5DRRA 4J3DA 1XXEA 4FW3A 4ISAA 4MQYA 3PS3A 4FW4A 5DRPA 4OKGA 5DRQA 4LCHA 3P3EA 4U3DA 4FW6A 3PS1A 2IESA 2VESA 2GO4A 3UHMA 4OZEA 2JT2A 1YHCA
chains in the Genus database with same CATH homology


 
#chains in the Genus database with same CATH superfamily
 3P76 A;  4LCF A;  2J65 A;  4IS9 A;  4LCG A;  4FW5 A;  2GO3 A;  1P42 A;  3PS2 A;  3P3C A;  4FW7 A;  5DRO A;  2O3Z A;  4U3B A;  3NZK A;  1YH8 A;  3U1Y A;  2IER A;  4MDT A;  3P3G A;  5DRR A;  4J3D A;  1XXE A;  4FW3 A;  4ISA A;  4MQY A;  3PS3 A;  4FW4 A;  5DRP A;  4OKG A;  5DRQ A;  4LCH A;  3P3E A;  4U3D A;  4FW6 A;  3PS1 A;  2IES A;  2VES A;  2GO4 A;  3UHM A;  4OZE A;  2JT2 A;  1YHC A; 
#chains in the Genus database with same CATH topology
 5JEA E;  1OYP A;  4JI7 I;  1KIJ A;  4FW5 A;  3T1H E;  5A2Q C;  2UUC E;  3K17 A;  2F4V I;  1R6M A;  2C38 B;  4DV4 E;  2UXC E;  4IFD A;  4HAC B;  1YH8 A;  1UDS A;  2NN6 E;  2ZM6 I;  1N36 I;  2UU9 I;  1XMO I;  2JEA B;  1S4E A;  5L3J A;  4KHP I;  3M85 D;  4QNJ A;  4LF5 E;  4DR6 I;  3F0N A;  4WUC A;  4JI5 E;  2PO1 A;  1ZM9 A;  2V8P A;  2VES A;  2C37 B;  4X66 I;  2OI2 A;  2IOQ A;  5IWA I;  1J5E E;  2C39 B;  1H7S A;  3ZKD A;  2Z5E A;  2HHH I;  2BA0 G;  4JI0 E;  2XEX A;  2PO2 A;  2V34 A;  2WW4 A;  4DV1 I;  4IZJ A;  4ED4 A;  2X36 A;  3Q6M A;  4JG4 A;  4RPF A;  4NXM E;  4P52 A;  4B3T I;  5K36 A;  2WNR B;  4DV6 I;  2UXD E;  1USU A;  2AJ4 A;  2JT2 A;  1MU5 A;  4DR2 I;  5C0X A;  1NHI A;  1H73 A;  5DRO A;  1HNX I;  4PRX A;  4DUY E;  3JAM C;  4WUD A;  1IBL I;  4DV2 E;  3U1Y A;  4UT4 A;  5DNX A;  4IFD B;  4DV3 I;  1IBK E;  4UTG A;  5FLX Q;  1N34 I;  2E5L E;  4MU3 A;  4DR3 E;  1Z0W A;  4I3H A;  1HNW I;  2Z5C B;  4JV5 I;  2BM1 A;  3OTO I;  4IFD C;  4B3S I;  4LF4 E;  2O1V A;  2PO1 B;  1XNR E;  2NN6 A;  1UDQ A;  1XHK A;  3U1K A;  4EMD A;  4OO1 F;  2GS8 A;  1FKA E;  4JI4 I;  3HUL A;  1VIS A;  2CG9 A;  4DV0 I;  3J81 C;  2O1T A;  4JYA I;  1HNZ E;  2PO2 B;  2AE8 A;  4FW9 A;  4LF9 E;  4X64 E;  5LMN I;  1PVG A;  4M1K A;  4MYT A;  1IBM I;  1ZM4 A;  4IZJ B;  4JI6 I;  4OX9 I;  3HKM A;  3D4J A;  4YY3 I;  1BKN A;  1ELO A;  4ZVI A;  5K36 B;  5JEA F;  5EL9 A;  2BA0 D;  4JI7 E;  4PU9 A;  4NBQ A;  2WP8 A;  3QT8 A;  3KRN A;  2PNL A;  5C0X B;  2F4V E;  1XNQ I;  3ZZT A;  5LMU I;  1N32 I;  3J80 Q;  2CVE A;  4JI1 I;  4GKJ I;  2ZM6 E;  3DD6 A;  1N36 E;  2UU9 E;  1XMO E;  4AIM A;  2E1R A;  4KHP E;  3H1C A;  4DR6 E;  2PO0 A;  2CGE A;  1KTV A;  4DPY A;  4X66 E;  2UXB I;  2VQF I;  3B8H A;  4IFD D;  2IES A;  5IWA E;  3GON A;  3B4T A;  4DV1 E;  4JI3 I;  2PNM A;  2NPF A;  4LF7 I;  4DV7 I;  2Z5B B;  3K85 A;  4AID A;  4URL A;  4DPW A;  5DRR A;  4B3T E;  4B3M I;  4DV6 E;  2DEI A;  4GKK I;  2R3V A;  5FWK A;  3Q6N A;  5LL6 R;  2O1W A;  1USV A;  4DR2 E;  4LFA I;  1NHJ A;  2WP8 B;  2HKJ A;  3PS2 A;  1IBL E;  4YHH I;  2BR2 A;  3P06 A;  2UUA E;  4DV3 E;  1I94 I;  4DPU A;  2HK3 A;  1N34 E;  1FWL A;  1HNW E;  5C0X D;  4JV5 E;  4OO1 A;  1ZM2 A;  2PO0 B;  4DR5 I;  1FNM A;  4FW6 A;  4JI4 E;  4GQU A;  4N3O A;  4MYU A;  2PNZ A;  1Z0C A;  2UUB I;  3P3C A;  2ZBK B;  1Z0V A;  4LFC E;  4DR7 I;  5LMN E;  4DR4 E;  1IBM E;  4LF8 I;  2GQ0 A;  5FLX C;  1Y60 A;  1XMQ E;  1KKH A;  2VF3 A;  4AQY I;  4JI6 E;  4OX9 E;  1Z5C A;  4YY3 E;  3CDI A;  1MG7 A;  5JEA A;  1Y6Z A;  4BA1 A;  2X7I A;  2V2Z A;  2JE6 A;  1N0V C;  4OZE A;  1XNQ E;  1ZM3 A;  1Z5B A;  5LMU E;  1N32 E;  3ZZU A;  1J5E I;  3M7N G;  4JI1 E;  4MU4 A;  4DPX A;  4GKJ E;  4DV5 I;  1HR0 E;  1WUU A;  1Z0T A;  1FJG I;  5IT9 Q;  3WU6 A;  5BR8 I;  2HFS A;  1H72 C;  3T1Y I;  4B3R I;  1KVK A;  3L7Z A;  4OO1 B;  3WU5 A;  2VQF E;  2EFG A;  4X65 I;  2F1D A;  4U3D A;  2O1U A;  1R6L A;  4IFD E;  3J7A G;  4HYM A;  4FWD A;  2CZ9 A;  1H7U A;  4JI3 E;  1MX0 A;  2PNZ B;  3J80 C;  3GLL A;  4LF7 E;  3PYD A;  5K36 C;  4DUZ I;  4DV7 E;  1FWK A;  1PIE A;  2O3Z A;  4B3M E;  4R1F A;  1B63 A;  5C0X C;  1HK7 A;  5JEA B;  4GKK E;  2V2V A;  4BA1 B;  3WU3 A;  2V2Q A;  4JI2 I;  2C37 A;  4LFA E;  2JE6 B;  4NXN I;  4LF6 I;  2NN6 B;  2HHH E;  1D6T A;  2A2D A;  2R42 A;  4WUB A;  4YHH E;  1P42 A;  2JEB A;  2BV3 A;  2J7K A;  1Z59 A;  1I94 E;  4DR1 I;  1Z0B A;  2NN6 C;  1HNZ I;  1K47 A;  3P3G A;  3GCM A;  4IZK A;  2VQE I;  4LF9 I;  4X64 I;  4LOM A;  2Q2E B;  3PYF A;  1N33 I;  3L7Z B;  2BA1 G;  4MQY A;  4XTJ A;  1E3P A;  3CWV A;  3M7N D;  4DU8 A;  4BA2 A;  4IZJ D;  4K0K I;  3H4L A;  2UUB E;  4LFB I;  1YHC A;  4MU1 A;  4OO1 D;  3LTO A;  4LCF A;  4DR7 E;  4IS9 A;  4HXZ A;  1HNX E;  4JI8 I;  4LF8 E;  1Y4U A;  5K36 D;  4DPT A;  3ZM7 A;  4AQY E;  3V5R A;  5A2Q Q;  3GME A;  4X62 I;  4MU0 A;  3ZKB A;  3J7A M;  1Z0G A;  3PYE A;  2Z5C A;  1Y5Y A;  3OTO E;  4B3S E;  4FW4 A;  1ZXM A;  3ZZ0 A;  1OJ4 A;  2IOP A;  2GO4 A;  1Z0E A;  3LPS A;  4AM3 A;  4DV0 E;  3LNU A;  5JEA D;  4DV5 E;  4JYA E;  3B82 A;  4FW7 A;  1FJG E;  2JEB B;  5BR8 E;  3T1Y E;  4PRV A;  4Z7Y A;  5ELW A;  3T1H I;  4B3R E;  2UUC I;  5GME A;  1RRE A;  2IER A;  2NN6 D;  4MDT A;  4DV4 I;  1OYR A;  4IZK B;  1UDN A;  2UXC I;  1UEK A;  1XXE A;  4X65 E;  4ISA A;  3QT5 A;  4IFD F;  1NZ0 A;  4LF5 I;  2LJP A;  4JI5 I;  1N0U A;  4HY1 A;  4BA2 B;  4DUZ E;  1EA6 A;  3P76 A;  3WU4 A;  2DEJ A;  2J65 A;  4LCG A;  4FWG A;  5IT9 C;  1ZXN A;  1DAR A;  1H74 A;  4JI0 I;  2BR2 B;  3M6A A;  2UUA I;  4USM A;  4USK A;  1FI4 A;  1OYS A;  3QT6 A;  2BA1 D;  3JAM Q;  3PRY A;  1B62 A;  4J3D A;  4JI2 E;  4LPF A;  5DRP A;  2UXB E;  4NXM I;  4LCH A;  4NXN E;  3PYG A;  4LF6 E;  3PS1 A;  3M85 G;  1Y4S A;  2UXD I;  4OO1 C;  5K36 F;  4DR1 E;  4Z7C A;  2C38 A;  2A2C A;  4KQV A;  1Z5A A;  2Z5B A;  2ZIT A;  4QNK A;  5EKW A;  4LFC I;  3J81 Q;  2VQE E;  1UDO A;  4HAC A;  4DXL A;  4DR4 I;  4U3B A;  3NZK A;  5C0X F;  4DUY I;  1RHY A;  1XMQ I;  3Q1Q A;  1N33 E;  4DV2 I;  3QT7 A;  2BM0 A;  2JEA A;  1E3H A;  1IBK I;  3PS3 A;  1PKP A;  1S16 A;  2E5L I;  4OKG A;  4DR3 I;  4K0K E;  3B78 A;  4LFB E;  4DU7 A;  4P7A A;  3UHM A;  4LF4 I;  5JEA C;  2C39 A;  3CDJ A;  1XNR I;  3K1J A;  4JI8 E;  4FWH A;  4OO1 E;  1U2R A;  5DNL A;  2GO3 A;  2HFU A;  2HKE A;  2NN6 F;  2HK2 A;  5K36 E;  1HR0 I;  4X62 E;  1QZR A;  1NHH A;  1EI1 A;  3HJC A;  4FW3 A;  1VI7 A;  5C0X E;  2WNR A;  4DR5 E;  5DRQ A;  3P3E A;  5GMD A;  1RR9 A;  1A6F A;  4FN5 A;  3V2U C; 
#chains in the Genus database with same CATH homology
 3P76 A;  4LCF A;  2J65 A;  4IS9 A;  4LCG A;  4FW5 A;  2GO3 A;  1P42 A;  3PS2 A;  3P3C A;  4FW7 A;  5DRO A;  2O3Z A;  4U3B A;  3NZK A;  1YH8 A;  3U1Y A;  2IER A;  4MDT A;  3P3G A;  5DRR A;  4J3D A;  1XXE A;  4FW3 A;  4ISA A;  4MQY A;  3PS3 A;  4FW4 A;  5DRP A;  4OKG A;  5DRQ A;  4LCH A;  3P3E A;  4U3D A;  4FW6 A;  3PS1 A;  2IES A;  2VES A;  2GO4 A;  3UHM A;  4OZE A;  2JT2 A;  1YHC A; 
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similar chains in the Genus database (?% sequence similarity)
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similar chains in the pdb database (?% sequence similarity)

 
#similar chains in the Genus database (?% sequence similarity)
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#similar chains, but unknotted
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#similar chains in the pdb database (?% sequence similarity)
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