1YHCA

Crystal structure of aquifex aeolicus lpxc deacetylase complexed with cacodylate
Total Genus 91
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The genus trace: a function that shows values of genus (vertical axis) for subchains spanned between the first residue, and all other residues (shown on horizontal axis). The number of the latter residue and the genus of a given subchain are shown interactively.

Total Genus
91
sequence length
266
structure length
266
Chain Sequence
GLEKTVKEKLSFEGVGIHTGEYSKLIIHPEKEGTGIRFFKNGVYIPARHEFVVHTNHSTDLGFKGQRIKTVEHILSVLHLLEITNVTIEVIGNEIPILDGSGWEFYEAIRKNILNQNREIDYFVVEEPIIVEDEGRLIKAEPSDTLEVTYEGEFKNFLGRQKFTFVEGNEEEIVLARTFAFDWEIEHIKKVGLGKGGSLKNTLVLGKDKVYNPEGLRYENEPVRHKVFDLIGDLYLLGSPVKGKFYSFRGGHSLNVKLVKELAKKQ
Loading diagram...

The genus matrix. At position (x,y) a genus value for a subchain spanned between x’th and y’th residue is shown. Values of the genus are represented by color, according to the scale given on the right.

Structure visualization

After clicking on a point (x,y) in the genus matrix above, a subchain from x to y is shown in color.

Chord Diagram
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publication title UDP-3-O-((R)-3-hydroxymyristoyl)-N-acetylglucosamine Deacetylase Functions through a General Acid-Base Catalyst Pair Mechanism
pubmed doi rcsb
source organism Aquifex aeolicus
molecule tags Hydrolase
molecule keywords UDP-3-O-[3-hydroxymyristoyl] N-acetylglucosamine deacetylase
total genus 91
structure length 266
sequence length 266
chains with identical sequence B
ec nomenclature ec 3.5.1.108: UDP-3-O-acyl-N-acetylglucosamine deacetylase.
pdb deposition date 2005-01-07

pfam database annotations

chain Pfam Accession Code Pfam Family Identifier Pfam Description
A PF03331 LpxC UDP-3-O-acyl N-acetylglycosamine deacetylase
Image from the rcsb pdb (www.rcsb.org)
cath code
ClassArchitectureTopologyHomologyDomain
3.30.230.20 Alpha Beta 2-Layer Sandwich Ribosomal Protein S5; domain 2 lpxc deacetylase, domain 1 1yhcA01
3.30.1700.10 Alpha Beta 2-Layer Sandwich lpxc deacetylase, domain 1 lpxc deacetylase, domain 2 1yhcA02
4OZEA 4U3DA 3U1YA 5DROA 2JT2A 3PS2A 3NZKA 5DRQA 4FW5A 2GO4A 4FW4A 4FW7A 4MDTA 1P42A 2J65A 4IS9A 3PS3A 2IESA 4LCFA 2IERA 3PS1A 4FW3A 4LCHA 4ISAA 4U3BA 4MQYA 5DRRA 1YHCA 4OKGA 3P3CA 3UHMA 3P76A 4J3DA 2GO3A 4LCGA 2VESA 1XXEA 2O3ZA 5DRPA 1YH8A 3P3EA 4FW6A 3P3GA
chains in the Genus database with same CATH superfamily
1UDSA 1Z0EA 4XTJA 3B82A 4DV0E 1XHKA 4UT4A 4JI6I 3PYFA 2WP8B 1D6TA 1FJGI 5EKWA 2PNMA 4JI7E 1FI4A 4AIDA 1VI7A 1HNXE 4FW3A 3GCMA 4Z7YA 4U3BA 1IBKI 4J3DA 2BR2B 2VF3A 2PO0B 4YHHE 4OX9I 4OO1D 1HNZE 4JG4A 1E3PA 4JV5I 5C0XB 1PKPA 1S4EA 4DV6I 3CDJA 4B3SI 5JEAC 4B3TI 4DV7E 4JYAE 1KTVA 4IFDF 5K36C 4LF5E 4DR4E 4BA1A 5C0XF 4LF5I 2R3VA 3B4TA 4LF4E 4RPFA 4DV0I 3GLLA 4WUDA 1HR0I 4DR5I 3KRNA 3QT8A 4LPFA 1XNRI 3GMEA 4HYMA 1NHJA 2BM0A 1RHYA 1Z5CA 3PS2A 4DXLA 1N34I 3B78A 3J80Q 4LF8E 3ZM7A 2HKJA 2E1RA 1R6LA 2A2CA 4DV3E 4JI4I 2HKEA 2ZM6I 4HY1A 3PS1A 2UU9I 4ISAA 5DRRA 2WNRB 4LFBI 5BR8E 4DV5E 3P76A 1N36I 1XMOI 4JI0E 3OTOI 2UUAI 4LF6I 4IFDA 4MYUA 2R42A 1NHHA 2X36A 4LF9I 4JI8I 4QNKA 4BA1B 3QT7A 2BA1G 2Q2EB 1IBLI 2BM1A 1XNRE 1Z0GA 4GQUA 2HHHE 1Y4SA 2HHHI 4DV1I 1OYPA 3QT5A 2V2VA 5L3JA 2IERA 4DV4I 2HK2A 1K47A 1ZM3A 2UUCI 1Z5AA 4DR6I 5K36D 4DUZE 1N33E 1B62A 2UXCE 4OO1B 5JEAF 2E5LI 3H1CA 3H4LA 4B3SE 4DV2E 2LJPA 1H7UA 4FW5A 4DUYI 1N0VC 1FNMA 2PNLA 4DPUA 2NN6E 4LF9E 4X62E 1A6FA 4AM3A 2F4VI 1N32E 3M7NG 3HJCA 4X62I 4DPTA 1IBLE 1XMQE 4DR5E 4IZJD 3LNUA 2UXDE 2JEAA 4NXNE 2AE8A 2C38A 4JI7I 4WUCA 4DV4E 4U3DA 4NXME 4GKJE 4GKJI 2ZITA 1QZRA 1Z0BA 3T1YE 4B3MI 4JI6E 4X65E 4GKKE 2VQFI 1N0UA 2DEIA 2UUBI 1Z0TA 3LTOA 3HKMA 4JV5E 1H7SA 5K36B 3DD6A 3CDIA 2PO1A 1IBKE 4LF6E 2NN6D 3V5RA 1IBMI 4JI3I 2PO1B 4OX9E 5C0XC 1FKAE 3M7ND 5GMEA 2CVEA 3P3GA 3PYDA 4OZEA 1HNWE 3J80C 2UU9E 4BA2B 1ZXMA 4HXZA 1Z0CA 4DV5I 5BR8I 2JE6A 2WP8A 3PYEA 3M85G 2IESA 4X66E 4NXMI 2CGEA 4HACB 3F0NA 4R1FA 2UUCE 3M6AA 1E3HA 4OKGA 4JI1E 5DNLA 2O3ZA 5A2QQ 1YH8A 4QNJA 2GS8A 1MU5A 3NZKA 4AIMA 3Q6MA 2NN6A 3ZKBA 3GONA 1HNXI 2WNRA 5JEAB 1XMOE 1KVKA 4MDTA 2Z5BB 4DR7E 5LMNI 3ZKDA 4JI4E 3WU3A 1BKNA 1USUA 1Y5YA 5LL6R 2AJ4A 4IZKA 4JYAI 4LCGA 1XXEA 4URLA 3U1YA 2V34A 5C0XE 2F4VE 2V2ZA 2UXBE 4JI8E 3T1HI 2CZ9A 4FWHA 2F1DA 3T1YI 4B3ME 1FJGE 3L7ZB 4GKKI 2JEBA 1P42A 4HACA 2OI2A 1B63A 4LCHA 3ZZ0A 4DPXA 1VISA 3J7AM 3Q1QA 5EL9A 4DR1E 2Z5EA 1EA6A 1ZM2A 4LF7E 1UEKA 4DR6E 4LF7I 4IFDB 3HULA 4MU3A 2V2QA 1HNWI 4B3RE 2ZM6E 5LMUI 3U1KA 1FWKA 4YY3I 2V8PA 3V2UC 4UTGA 4LOMA 2PO2A 1IBME 4JI3E 3PS3A 3D4JA 4NBQA 2NN6C 4MU1A 2PO2B 2UXBI 1KKHA 1WUUA 4LFCI 4M1KA 1HR0E 4DR2E 3WU5A 2O1WA 4I3HA 5IT9Q 2VQEI 3JAMC 1PVGA 1DARA 4JI2E 1Z0VA 1R6MA 4NXNI 2Z5CA 5IWAE 3J81C 3P3EA 4DU7A 5IWAI 5DROA 1Z0WA 5DRQA 4DR1I 1ELOA 2WW4A 2BV3A 3L7ZA 4LF8I 1H73A 5FWKA 4OO1F 1NHIA 2EFGA 5K36F 5LMNE 1HNZI 4MU0A 4X64I 4DR3E 4DV3I 2UUBE 2C39B 3LPSA 5C0XA 2PO0A 3P3CA 4IFDD 2GO3A 3OTOE 2UUAE 2JE6B 1N36E 4P7AA 5FLXC 5A2QC 2C37A 5C0XD 2O1TA 4FWGA 4JI5E 2GQ0A 4FN5A 3CWVA 4AQYE 5GMDA 4K0KI 1UDQA 4PRVA 4DV1E 1USVA 2HK3A 1UDOA 2Z5CB 5DNXA 1U2RA 1UDNA 1MG7A 3T1HE 2BA1D 5JEAA 2JEBB 1PIEA 4ZVIA 2HFUA 5LMUE 4KQVA 3K85A 1Z5BA 1KIJA 4DV2I 5ELWA 1EI1A 1ZM4A 4OO1A 4YY3E 2IOPA 4LFBE 4FW4A 4DR4I 4MU4A 2IOQA 2HFSA 3Q6NA 1Y60A 3ZZTA 1NZ0A 1HK7A 4IZJA 5K36E 1FWLA 1S16A 2DEJA 4MQYA 3PRYA 2NPFA 2UXDI 5JEAE 3UHMA 4B3TE 2BA0D 4DPYA 4N3OA 2C37B 5DRPA 4WUBA 2JEAB 3WU6A 3J7AG 1MX0A 4P52A 1ZXNA 3P06A 2O1VA 2PNZA 2VQFE 4PRXA 4JI1I 1RREA 2PNZB 4IS9A 4USKA 4X65I 4B3RI 5K36A 2E5LE 3WU4A 5JEAD 1H72C 2J7KA 4DV7I 3K1JA 1J5EE 1J5EI 5IT9C 3JAMQ 3J81Q 4DR7I 2X7IA 4LFCE 1XMQI 1OYSA 4BA2A 4ED4A 1OJ4A 4DR2I 4IZKB 1Y4UA 1Y6ZA 4DV6E 2BA0G 4LFAE 3K17A 1H74A 4IFDE 4FW9A 4OO1C 4X66I 4IZJB 4LF4I 4USMA 3ZZUA 1YHCA 2ZBKB 4Z7CA 2CG9A 2O1UA 4DUZI 2A2DA 4AQYI 1N33I 2UXCI 2VQEE 5FLXQ 4K0KE 4DPWA 2C38B 4DU8A 2NN6B 4JI2I 4FW6A 2XEXA 3M85D 2JT2A 2BR2A 1N34E 3B8HA 1ZM9A 2C39A 2GO4A 2NN6F 3QT6A 4FW7A 4YHHI 1I94E 1XNQE 1OYRA 4MYTA 2J65A 1I94I 1XNQI 3PYGA 4JI5I 4OO1E 1N32I 4EMDA 4X64E 4DR3I 4LCFA 2Z5BA 4IFDC 4FWDA 4PU9A 4KHPI 4KHPE 4LFAI 4DUYE 2VESA 4JI0I 1RR9A 1Z59A
chains in the Genus database with same CATH topology
4OZEA 4U3DA 3U1YA 5DROA 2JT2A 3PS2A 3NZKA 5DRQA 4FW5A 2GO4A 4FW4A 4FW7A 4MDTA 1P42A 2J65A 4IS9A 3PS3A 2IESA 4LCFA 2IERA 3PS1A 4FW3A 4LCHA 4ISAA 4U3BA 4MQYA 5DRRA 1YHCA 4OKGA 3P3CA 3UHMA 3P76A 4J3DA 2GO3A 4LCGA 2VESA 1XXEA 2O3ZA 5DRPA 1YH8A 3P3EA 4FW6A 3P3GA
chains in the Genus database with same CATH homology


 
#chains in the Genus database with same CATH superfamily
 4OZE A;  4U3D A;  3U1Y A;  5DRO A;  2JT2 A;  3PS2 A;  3NZK A;  5DRQ A;  4FW5 A;  2GO4 A;  4FW4 A;  4FW7 A;  4MDT A;  1P42 A;  2J65 A;  4IS9 A;  3PS3 A;  2IES A;  4LCF A;  2IER A;  3PS1 A;  4FW3 A;  4LCH A;  4ISA A;  4U3B A;  4MQY A;  5DRR A;  1YHC A;  4OKG A;  3P3C A;  3UHM A;  3P76 A;  4J3D A;  2GO3 A;  4LCG A;  2VES A;  1XXE A;  2O3Z A;  5DRP A;  1YH8 A;  3P3E A;  4FW6 A;  3P3G A; 
#chains in the Genus database with same CATH topology
 1UDS A;  1Z0E A;  4XTJ A;  3B82 A;  4DV0 E;  1XHK A;  4UT4 A;  4JI6 I;  3PYF A;  2WP8 B;  1D6T A;  1FJG I;  5EKW A;  2PNM A;  4JI7 E;  1FI4 A;  4AID A;  1VI7 A;  1HNX E;  4FW3 A;  3GCM A;  4Z7Y A;  4U3B A;  1IBK I;  4J3D A;  2BR2 B;  2VF3 A;  2PO0 B;  4YHH E;  4OX9 I;  4OO1 D;  1HNZ E;  4JG4 A;  1E3P A;  4JV5 I;  5C0X B;  1PKP A;  1S4E A;  4DV6 I;  3CDJ A;  4B3S I;  5JEA C;  4B3T I;  4DV7 E;  4JYA E;  1KTV A;  4IFD F;  5K36 C;  4LF5 E;  4DR4 E;  4BA1 A;  5C0X F;  4LF5 I;  2R3V A;  3B4T A;  4LF4 E;  4RPF A;  4DV0 I;  3GLL A;  4WUD A;  1HR0 I;  4DR5 I;  3KRN A;  3QT8 A;  4LPF A;  1XNR I;  3GME A;  4HYM A;  1NHJ A;  2BM0 A;  1RHY A;  1Z5C A;  3PS2 A;  4DXL A;  1N34 I;  3B78 A;  3J80 Q;  4LF8 E;  3ZM7 A;  2HKJ A;  2E1R A;  1R6L A;  2A2C A;  4DV3 E;  4JI4 I;  2HKE A;  2ZM6 I;  4HY1 A;  3PS1 A;  2UU9 I;  4ISA A;  5DRR A;  2WNR B;  4LFB I;  5BR8 E;  4DV5 E;  3P76 A;  1N36 I;  1XMO I;  4JI0 E;  3OTO I;  2UUA I;  4LF6 I;  4IFD A;  4MYU A;  2R42 A;  1NHH A;  2X36 A;  4LF9 I;  4JI8 I;  4QNK A;  4BA1 B;  3QT7 A;  2BA1 G;  2Q2E B;  1IBL I;  2BM1 A;  1XNR E;  1Z0G A;  4GQU A;  2HHH E;  1Y4S A;  2HHH I;  4DV1 I;  1OYP A;  3QT5 A;  2V2V A;  5L3J A;  2IER A;  4DV4 I;  2HK2 A;  1K47 A;  1ZM3 A;  2UUC I;  1Z5A A;  4DR6 I;  5K36 D;  4DUZ E;  1N33 E;  1B62 A;  2UXC E;  4OO1 B;  5JEA F;  2E5L I;  3H1C A;  3H4L A;  4B3S E;  4DV2 E;  2LJP A;  1H7U A;  4FW5 A;  4DUY I;  1N0V C;  1FNM A;  2PNL A;  4DPU A;  2NN6 E;  4LF9 E;  4X62 E;  1A6F A;  4AM3 A;  2F4V I;  1N32 E;  3M7N G;  3HJC A;  4X62 I;  4DPT A;  1IBL E;  1XMQ E;  4DR5 E;  4IZJ D;  3LNU A;  2UXD E;  2JEA A;  4NXN E;  2AE8 A;  2C38 A;  4JI7 I;  4WUC A;  4DV4 E;  4U3D A;  4NXM E;  4GKJ E;  4GKJ I;  2ZIT A;  1QZR A;  1Z0B A;  3T1Y E;  4B3M I;  4JI6 E;  4X65 E;  4GKK E;  2VQF I;  1N0U A;  2DEI A;  2UUB I;  1Z0T A;  3LTO A;  3HKM A;  4JV5 E;  1H7S A;  5K36 B;  3DD6 A;  3CDI A;  2PO1 A;  1IBK E;  4LF6 E;  2NN6 D;  3V5R A;  1IBM I;  4JI3 I;  2PO1 B;  4OX9 E;  5C0X C;  1FKA E;  3M7N D;  5GME A;  2CVE A;  3P3G A;  3PYD A;  4OZE A;  1HNW E;  3J80 C;  2UU9 E;  4BA2 B;  1ZXM A;  4HXZ A;  1Z0C A;  4DV5 I;  5BR8 I;  2JE6 A;  2WP8 A;  3PYE A;  3M85 G;  2IES A;  4X66 E;  4NXM I;  2CGE A;  4HAC B;  3F0N A;  4R1F A;  2UUC E;  3M6A A;  1E3H A;  4OKG A;  4JI1 E;  5DNL A;  2O3Z A;  5A2Q Q;  1YH8 A;  4QNJ A;  2GS8 A;  1MU5 A;  3NZK A;  4AIM A;  3Q6M A;  2NN6 A;  3ZKB A;  3GON A;  1HNX I;  2WNR A;  5JEA B;  1XMO E;  1KVK A;  4MDT A;  2Z5B B;  4DR7 E;  5LMN I;  3ZKD A;  4JI4 E;  3WU3 A;  1BKN A;  1USU A;  1Y5Y A;  5LL6 R;  2AJ4 A;  4IZK A;  4JYA I;  4LCG A;  1XXE A;  4URL A;  3U1Y A;  2V34 A;  5C0X E;  2F4V E;  2V2Z A;  2UXB E;  4JI8 E;  3T1H I;  2CZ9 A;  4FWH A;  2F1D A;  3T1Y I;  4B3M E;  1FJG E;  3L7Z B;  4GKK I;  2JEB A;  1P42 A;  4HAC A;  2OI2 A;  1B63 A;  4LCH A;  3ZZ0 A;  4DPX A;  1VIS A;  3J7A M;  3Q1Q A;  5EL9 A;  4DR1 E;  2Z5E A;  1EA6 A;  1ZM2 A;  4LF7 E;  1UEK A;  4DR6 E;  4LF7 I;  4IFD B;  3HUL A;  4MU3 A;  2V2Q A;  1HNW I;  4B3R E;  2ZM6 E;  5LMU I;  3U1K A;  1FWK A;  4YY3 I;  2V8P A;  3V2U C;  4UTG A;  4LOM A;  2PO2 A;  1IBM E;  4JI3 E;  3PS3 A;  3D4J A;  4NBQ A;  2NN6 C;  4MU1 A;  2PO2 B;  2UXB I;  1KKH A;  1WUU A;  4LFC I;  4M1K A;  1HR0 E;  4DR2 E;  3WU5 A;  2O1W A;  4I3H A;  5IT9 Q;  2VQE I;  3JAM C;  1PVG A;  1DAR A;  4JI2 E;  1Z0V A;  1R6M A;  4NXN I;  2Z5C A;  5IWA E;  3J81 C;  3P3E A;  4DU7 A;  5IWA I;  5DRO A;  1Z0W A;  5DRQ A;  4DR1 I;  1ELO A;  2WW4 A;  2BV3 A;  3L7Z A;  4LF8 I;  1H73 A;  5FWK A;  4OO1 F;  1NHI A;  2EFG A;  5K36 F;  5LMN E;  1HNZ I;  4MU0 A;  4X64 I;  4DR3 E;  4DV3 I;  2UUB E;  2C39 B;  3LPS A;  5C0X A;  2PO0 A;  3P3C A;  4IFD D;  2GO3 A;  3OTO E;  2UUA E;  2JE6 B;  1N36 E;  4P7A A;  5FLX C;  5A2Q C;  2C37 A;  5C0X D;  2O1T A;  4FWG A;  4JI5 E;  2GQ0 A;  4FN5 A;  3CWV A;  4AQY E;  5GMD A;  4K0K I;  1UDQ A;  4PRV A;  4DV1 E;  1USV A;  2HK3 A;  1UDO A;  2Z5C B;  5DNX A;  1U2R A;  1UDN A;  1MG7 A;  3T1H E;  2BA1 D;  5JEA A;  2JEB B;  1PIE A;  4ZVI A;  2HFU A;  5LMU E;  4KQV A;  3K85 A;  1Z5B A;  1KIJ A;  4DV2 I;  5ELW A;  1EI1 A;  1ZM4 A;  4OO1 A;  4YY3 E;  2IOP A;  4LFB E;  4FW4 A;  4DR4 I;  4MU4 A;  2IOQ A;  2HFS A;  3Q6N A;  1Y60 A;  3ZZT A;  1NZ0 A;  1HK7 A;  4IZJ A;  5K36 E;  1FWL A;  1S16 A;  2DEJ A;  4MQY A;  3PRY A;  2NPF A;  2UXD I;  5JEA E;  3UHM A;  4B3T E;  2BA0 D;  4DPY A;  4N3O A;  2C37 B;  5DRP A;  4WUB A;  2JEA B;  3WU6 A;  3J7A G;  1MX0 A;  4P52 A;  1ZXN A;  3P06 A;  2O1V A;  2PNZ A;  2VQF E;  4PRX A;  4JI1 I;  1RRE A;  2PNZ B;  4IS9 A;  4USK A;  4X65 I;  4B3R I;  5K36 A;  2E5L E;  3WU4 A;  5JEA D;  1H72 C;  2J7K A;  4DV7 I;  3K1J A;  1J5E E;  1J5E I;  5IT9 C;  3JAM Q;  3J81 Q;  4DR7 I;  2X7I A;  4LFC E;  1XMQ I;  1OYS A;  4BA2 A;  4ED4 A;  1OJ4 A;  4DR2 I;  4IZK B;  1Y4U A;  1Y6Z A;  4DV6 E;  2BA0 G;  4LFA E;  3K17 A;  1H74 A;  4IFD E;  4FW9 A;  4OO1 C;  4X66 I;  4IZJ B;  4LF4 I;  4USM A;  3ZZU A;  1YHC A;  2ZBK B;  4Z7C A;  2CG9 A;  2O1U A;  4DUZ I;  2A2D A;  4AQY I;  1N33 I;  2UXC I;  2VQE E;  5FLX Q;  4K0K E;  4DPW A;  2C38 B;  4DU8 A;  2NN6 B;  4JI2 I;  4FW6 A;  2XEX A;  3M85 D;  2JT2 A;  2BR2 A;  1N34 E;  3B8H A;  1ZM9 A;  2C39 A;  2GO4 A;  2NN6 F;  3QT6 A;  4FW7 A;  4YHH I;  1I94 E;  1XNQ E;  1OYR A;  4MYT A;  2J65 A;  1I94 I;  1XNQ I;  3PYG A;  4JI5 I;  4OO1 E;  1N32 I;  4EMD A;  4X64 E;  4DR3 I;  4LCF A;  2Z5B A;  4IFD C;  4FWD A;  4PU9 A;  4KHP I;  4KHP E;  4LFA I;  4DUY E;  2VES A;  4JI0 I;  1RR9 A;  1Z59 A; 
#chains in the Genus database with same CATH homology
 4OZE A;  4U3D A;  3U1Y A;  5DRO A;  2JT2 A;  3PS2 A;  3NZK A;  5DRQ A;  4FW5 A;  2GO4 A;  4FW4 A;  4FW7 A;  4MDT A;  1P42 A;  2J65 A;  4IS9 A;  3PS3 A;  2IES A;  4LCF A;  2IER A;  3PS1 A;  4FW3 A;  4LCH A;  4ISA A;  4U3B A;  4MQY A;  5DRR A;  1YHC A;  4OKG A;  3P3C A;  3UHM A;  3P76 A;  4J3D A;  2GO3 A;  4LCG A;  2VES A;  1XXE A;  2O3Z A;  5DRP A;  1YH8 A;  3P3E A;  4FW6 A;  3P3G A; 
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similar chains in the Genus database (?% sequence similarity)
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similar chains in the pdb database (?% sequence similarity)

 
#similar chains in the Genus database (?% sequence similarity)
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#similar chains, but unknotted
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#similar chains in the pdb database (?% sequence similarity)
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