1YHCA

Crystal structure of aquifex aeolicus lpxc deacetylase complexed with cacodylate
Total Genus 91
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The genus trace: a function that shows values of genus (vertical axis) for subchains spanned between the first residue, and all other residues (shown on horizontal axis). The number of the latter residue and the genus of a given subchain are shown interactively.

Total Genus
91
sequence length
266
structure length
266
Chain Sequence
GLEKTVKEKLSFEGVGIHTGEYSKLIIHPEKEGTGIRFFKNGVYIPARHEFVVHTNHSTDLGFKGQRIKTVEHILSVLHLLEITNVTIEVIGNEIPILDGSGWEFYEAIRKNILNQNREIDYFVVEEPIIVEDEGRLIKAEPSDTLEVTYEGEFKNFLGRQKFTFVEGNEEEIVLARTFAFDWEIEHIKKVGLGKGGSLKNTLVLGKDKVYNPEGLRYENEPVRHKVFDLIGDLYLLGSPVKGKFYSFRGGHSLNVKLVKELAKKQ
Loading diagram...

The genus matrix. At position (x,y) a genus value for a subchain spanned between x’th and y’th residue is shown. Values of the genus are represented by color, according to the scale given on the right.

Structure visualization

After clicking on a point (x,y) in the genus matrix above, a subchain from x to y is shown in color.

Chord Diagram
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publication title UDP-3-O-((R)-3-hydroxymyristoyl)-N-acetylglucosamine Deacetylase Functions through a General Acid-Base Catalyst Pair Mechanism
pubmed doi rcsb
source organism Aquifex aeolicus
molecule tags Hydrolase
molecule keywords UDP-3-O-[3-hydroxymyristoyl] N-acetylglucosamine deacetylase
total genus 91
structure length 266
sequence length 266
chains with identical sequence B
ec nomenclature ec 3.5.1.108: UDP-3-O-acyl-N-acetylglucosamine deacetylase.
pdb deposition date 2005-01-07

pfam database annotations

chain Pfam Accession Code Pfam Family Identifier Pfam Description
A PF03331 LpxC UDP-3-O-acyl N-acetylglycosamine deacetylase
Image from the rcsb pdb (www.rcsb.org)
cath code
ClassArchitectureTopologyHomologyDomain
3.30.230.20 Alpha Beta 2-Layer Sandwich Ribosomal Protein S5; domain 2 lpxc deacetylase, domain 1 1yhcA01
3.30.1700.10 Alpha Beta 2-Layer Sandwich lpxc deacetylase, domain 1 lpxc deacetylase, domain 2 1yhcA02
2J65A 4J3DA 4FW6A 5DRQA 5DRPA 3P3EA 3U1YA 3P3CA 2O3ZA 3PS2A 3PS1A 4LCHA 5DROA 3NZKA 4ISAA 2GO4A 4MDTA 1XXEA 1YHCA 4FW4A 4U3BA 3PS3A 2IESA 4U3DA 4MQYA 4FW3A 4OKGA 4FW7A 3P3GA 1YH8A 2GO3A 4IS9A 4LCFA 5DRRA 4OZEA 3UHMA 2IERA 4FW5A 2VESA 4LCGA 1P42A 2JT2A 3P76A
chains in the Genus database with same CATH superfamily
4JI7I 4HY1A 4JV5I 2HFUA 4LF6E 4OO1F 4P52A 5LMUI 5JEAD 4BA2B 2UUCE 1S16A 5JEAE 2E5LI 4FW9A 5A2QQ 4GKJE 2O1UA 5A2QC 1UDOA 5JEAF 4U3BA 3PS3A 4IFDD 3PYDA 4MQYA 1IBKE 4OO1E 5BR8E 1XMOE 4HACB 4IS9A 2VQFI 4OO1C 1Y6ZA 4LCFA 4DR7E 4LF8E 4JI0I 3D4JA 4DV3I 3JAMC 3PYFA 4DUZI 4K0KI 4YHHE 3U1YA 2AJ4A 4DR5E 4LPFA 2X7IA 2V2ZA 2HK2A 4DR5I 1RR9A 4LFAE 1H72C 1Z0WA 2IESA 1Y4SA 4DU7A 1Z0VA 5LL6R 3J80Q 2NN6E 4PU9A 1NHJA 2XEXA 1Y60A 3DD6A 2C38B 2PNZB 3L7ZA 5IT9C 1HNXE 4JI3E 3F0NA 4DPUA 3OTOI 5DNLA 2BR2A 4EMDA 4LF8I 5LMNE 4JI1I 5BR8I 1E3HA 5DROA 1YHCA 2UUBE 3K85A 4WUDA 4LF5I 1Z0CA 1H7SA 1N36I 2BM1A 2V34A 2WW4A 4ED4A 1HNZI 4LF6I 4MU0A 3ZKBA 4M1KA 4PRXA 4KHPE 2GS8A 4OO1A 2Z5CB 2Q2EB 4DXLA 3B8HA 4IFDB 1HR0I 1OYSA 4BA1A 1MG7A 4HXZA 1IBMI 4IFDF 2O1WA 2HHHE 5JEAA 1I94I 3QT5A 4AM3A 5IWAE 2UXCE 4Z7YA 4DR1E 4DV5E 1ZXNA 2HK3A 5ELWA 2HHHI 4JI8I 2DEIA 3M7NG 4LF4I 4NXNI 2BA1D 4LF5E 4JI6I 2VQEI 4LF9I 3HULA 1N36E 4X62I 1MU5A 3M85D 2UUBI 4B3RE 4OKGA 2PO2A 3Q1QA 3WU5A 1K47A 1YH8A 2PO0A 1N34E 2UUAE 2F4VE 2IOPA 2ZM6E 1QZRA 3CDJA 2NN6B 4BA2A 2WNRB 4DV6E 1B63A 4LFCI 1UDQA 4IZKB 4AIMA 1RREA 1Z5BA 5FWKA 4XTJA 5IT9Q 4FN5A 4DPWA 2GO4A 1XXEA 3T1YE 1PVGA 5C0XA 2UXDE 3LNUA 1H7UA 1DARA 4X66E 4JI2I 1S4EA 2V8PA 4N3OA 5K36F 1WUUA 3PYEA 3H1CA 3GCMA 1U2RA 4JYAE 4IFDE 4WUCA 4LCGA 4DV3E 2JT2A 4NBQA 3QT7A 2F1DA 1EI1A 2VF3A 4IFDA 2WP8A 3P3EA 4FWGA 1BKNA 1IBLI 1E3PA 1PKPA 2Z5BA 4FWDA 2ZBKB 4USKA 1B62A 3WU6A 5FLXC 2PO1A 4U3DA 1FJGI 2BA1G 4AIDA 4X62E 4LOMA 3T1HI 5EL9A 4JI6E 2VESA 1ZM4A 4DV4E 1D6TA 3U1KA 5L3JA 4DR1I 3CDIA 5DRPA 1Z0EA 4UTGA 2O3ZA 3PS2A 2JEAB 2J7KA 4YY3E 3ZZTA 1Z5CA 2C37B 1N33I 4NXME 4DV5I 4LFBI 1IBME 4MDTA 1Z0TA 2X36A 4MU3A 2Z5CA 4USMA 4OX9E 3ZZ0A 4HACA 4YY3I 2PO2B 2BA0D 3LPSA 3GLLA 1XNQI 2UU9E 3T1HE 5FLXQ 4B3SI 1HNZE 2V2VA 1J5EE 4IZJB 2LJPA 1RHYA 1N0UA 2UXBI 3GMEA 2ZITA 4B3TE 4LF7I 3K1JA 1Z0GA 1OJ4A 5C0XF 1FI4A 5K36D 1ELOA 5LMNI 2BM0A 1N32I 4UT4A 3OTOE 2AE8A 4DU8A 4FW3A 1R6LA 3ZM7A 4MYTA 2C39B 2V2QA 4JI4I 3HKMA 1Z0BA 2C38A 2PNZA 3UHMA 1R6MA 5GMDA 5JEAC 1Z5AA 1USVA 4LFAI 5EKWA 3WU4A 2E1RA 3P06A 1OYRA 4JYAI 2BA0G 3NZKA 1HNWE 2UXDI 4AQYE 4DUZE 1HNWI 2R3VA 2Z5EA 1ZM9A 4DR7I 2ZM6I 4YHHI 4FW5A 4K0KE 4B3ME 4MU1A 3B82A 5C0XB 2J65A 2JEBA 2DEJA 1MX0A 3J7AG 3HJCA 2F4VI 4GKJI 4JI4E 2VQFE 2JE6B 2NN6A 2UUCI 1Z59A 4LCHA 1NHHA 3PYGA 1UDSA 2A2DA 1ZM3A 5LMUE 3M7ND 3PRYA 4JI7E 4JV5E 4DV1E 2GO3A 4P7AA 1USUA 1XMQI 2UXBE 2NPFA 2EFGA 4NXNE 2CG9A 4OX9I 2UXCI 5IWAI 1J5EI 1KVKA 1ZM2A 2BR2B 1FWKA 4JI8E 4LFBE 1XNQE 5K36C 3H4LA 2HFSA 1UDNA 5K36A 1XNRI 2A2CA 3M6AA 4DR2I 4FW7A 2PNLA 4DR6E 2PO0B 4GQUA 2C39A 4IZJD 1FNMA 3LTOA 2R42A 1N0VC 1UEKA 1VISA 1PIEA 5DNXA 4DV0I 2PO1B 1KIJA 4OO1B 3Q6MA 2CZ9A 4J3DA 3P76A 4DUYI 4IZKA 4I3HA 1XMQE 4X65I 2HKEA 1FJGE 4JI1E 1FWLA 1KKHA 5C0XE 3L7ZB 4JI3I 4JI5E 5C0XD 3P3GA 1IBLE 2C37A 4OZEA 2BV3A 1NHIA 4DPXA 4HYMA 4GKKE 1HNXI 1P42A 4R1FA 3V5RA 1XHKA 3B4TA 4B3MI 4QNKA 4X66I 4JI2E 2JEBB 2PNMA 2NN6F 3T1YI 1FKAE 2O1VA 3PS1A 4IZJA 1NZ0A 2NN6D 3B78A 4FW4A 3KRNA 4AQYI 4DPYA 3ZKDA 3J80C 4GKKI 4IFDC 4JI0E 4RPFA 3QT6A 3K17A 3WU3A 2IERA 3QT8A 1XMOI 4LFCE 3J81C 2CVEA 4KQVA 1Y5YA 4DV6I 1IBKI 4B3RI 4JI5I 1EA6A 4FW6A 3M85G 1ZXMA 5K36E 1N32E 1H74A 2JEAA 1N34I 2E5LE 2UUAI 4LF9E 4PRVA 4B3TI 4LF7E 4ISAA 2VQEE 4FWHA 3CWVA 5K36B 4QNJA 4URLA 4X64I 3J7AM 4BA1B 4LF4E 2GQ0A 2IOQA 4DPTA 4DV2E 2HKJA 4MYUA 5JEAB 4X65E 4B3SE 4DR4I 3J81Q 2O1TA 4ZVIA 4DUYE 1KTVA 2NN6C 5GMEA 4DR3E 4DV0E 1VI7A 4JG4A 1XNRE 5DRQA 1A6FA 2WP8B 2CGEA 4DV7E 1HK7A 3P3CA 2UU9I 4DR2E 4DR6I 2Z5BB 3V2UC 3Q6NA 2WNRA 4MU4A 3GONA 3ZZUA 2OI2A 4NXMI 1I94E 4DR3I 4DV4I 1N33E 4KHPI 3JAMQ 2JE6A 5C0XC 5DRRA 1OYPA 4DR4E 4DV7I 1Y4UA 4DV2I 1HR0E 4OO1D 4Z7CA 4DV1I 1H73A 4WUBA 4X64E
chains in the Genus database with same CATH topology
2J65A 4J3DA 4FW6A 5DRQA 5DRPA 3P3EA 3U1YA 3P3CA 2O3ZA 3PS2A 3PS1A 4LCHA 5DROA 3NZKA 4ISAA 2GO4A 4MDTA 1XXEA 1YHCA 4FW4A 4U3BA 3PS3A 2IESA 4U3DA 4MQYA 4FW3A 4OKGA 4FW7A 3P3GA 1YH8A 2GO3A 4IS9A 4LCFA 5DRRA 4OZEA 3UHMA 2IERA 4FW5A 2VESA 4LCGA 1P42A 2JT2A 3P76A
chains in the Genus database with same CATH homology


 
#chains in the Genus database with same CATH superfamily
 2J65 A;  4J3D A;  4FW6 A;  5DRQ A;  5DRP A;  3P3E A;  3U1Y A;  3P3C A;  2O3Z A;  3PS2 A;  3PS1 A;  4LCH A;  5DRO A;  3NZK A;  4ISA A;  2GO4 A;  4MDT A;  1XXE A;  1YHC A;  4FW4 A;  4U3B A;  3PS3 A;  2IES A;  4U3D A;  4MQY A;  4FW3 A;  4OKG A;  4FW7 A;  3P3G A;  1YH8 A;  2GO3 A;  4IS9 A;  4LCF A;  5DRR A;  4OZE A;  3UHM A;  2IER A;  4FW5 A;  2VES A;  4LCG A;  1P42 A;  2JT2 A;  3P76 A; 
#chains in the Genus database with same CATH topology
 4JI7 I;  4HY1 A;  4JV5 I;  2HFU A;  4LF6 E;  4OO1 F;  4P52 A;  5LMU I;  5JEA D;  4BA2 B;  2UUC E;  1S16 A;  5JEA E;  2E5L I;  4FW9 A;  5A2Q Q;  4GKJ E;  2O1U A;  5A2Q C;  1UDO A;  5JEA F;  4U3B A;  3PS3 A;  4IFD D;  3PYD A;  4MQY A;  1IBK E;  4OO1 E;  5BR8 E;  1XMO E;  4HAC B;  4IS9 A;  2VQF I;  4OO1 C;  1Y6Z A;  4LCF A;  4DR7 E;  4LF8 E;  4JI0 I;  3D4J A;  4DV3 I;  3JAM C;  3PYF A;  4DUZ I;  4K0K I;  4YHH E;  3U1Y A;  2AJ4 A;  4DR5 E;  4LPF A;  2X7I A;  2V2Z A;  2HK2 A;  4DR5 I;  1RR9 A;  4LFA E;  1H72 C;  1Z0W A;  2IES A;  1Y4S A;  4DU7 A;  1Z0V A;  5LL6 R;  3J80 Q;  2NN6 E;  4PU9 A;  1NHJ A;  2XEX A;  1Y60 A;  3DD6 A;  2C38 B;  2PNZ B;  3L7Z A;  5IT9 C;  1HNX E;  4JI3 E;  3F0N A;  4DPU A;  3OTO I;  5DNL A;  2BR2 A;  4EMD A;  4LF8 I;  5LMN E;  4JI1 I;  5BR8 I;  1E3H A;  5DRO A;  1YHC A;  2UUB E;  3K85 A;  4WUD A;  4LF5 I;  1Z0C A;  1H7S A;  1N36 I;  2BM1 A;  2V34 A;  2WW4 A;  4ED4 A;  1HNZ I;  4LF6 I;  4MU0 A;  3ZKB A;  4M1K A;  4PRX A;  4KHP E;  2GS8 A;  4OO1 A;  2Z5C B;  2Q2E B;  4DXL A;  3B8H A;  4IFD B;  1HR0 I;  1OYS A;  4BA1 A;  1MG7 A;  4HXZ A;  1IBM I;  4IFD F;  2O1W A;  2HHH E;  5JEA A;  1I94 I;  3QT5 A;  4AM3 A;  5IWA E;  2UXC E;  4Z7Y A;  4DR1 E;  4DV5 E;  1ZXN A;  2HK3 A;  5ELW A;  2HHH I;  4JI8 I;  2DEI A;  3M7N G;  4LF4 I;  4NXN I;  2BA1 D;  4LF5 E;  4JI6 I;  2VQE I;  4LF9 I;  3HUL A;  1N36 E;  4X62 I;  1MU5 A;  3M85 D;  2UUB I;  4B3R E;  4OKG A;  2PO2 A;  3Q1Q A;  3WU5 A;  1K47 A;  1YH8 A;  2PO0 A;  1N34 E;  2UUA E;  2F4V E;  2IOP A;  2ZM6 E;  1QZR A;  3CDJ A;  2NN6 B;  4BA2 A;  2WNR B;  4DV6 E;  1B63 A;  4LFC I;  1UDQ A;  4IZK B;  4AIM A;  1RRE A;  1Z5B A;  5FWK A;  4XTJ A;  5IT9 Q;  4FN5 A;  4DPW A;  2GO4 A;  1XXE A;  3T1Y E;  1PVG A;  5C0X A;  2UXD E;  3LNU A;  1H7U A;  1DAR A;  4X66 E;  4JI2 I;  1S4E A;  2V8P A;  4N3O A;  5K36 F;  1WUU A;  3PYE A;  3H1C A;  3GCM A;  1U2R A;  4JYA E;  4IFD E;  4WUC A;  4LCG A;  4DV3 E;  2JT2 A;  4NBQ A;  3QT7 A;  2F1D A;  1EI1 A;  2VF3 A;  4IFD A;  2WP8 A;  3P3E A;  4FWG A;  1BKN A;  1IBL I;  1E3P A;  1PKP A;  2Z5B A;  4FWD A;  2ZBK B;  4USK A;  1B62 A;  3WU6 A;  5FLX C;  2PO1 A;  4U3D A;  1FJG I;  2BA1 G;  4AID A;  4X62 E;  4LOM A;  3T1H I;  5EL9 A;  4JI6 E;  2VES A;  1ZM4 A;  4DV4 E;  1D6T A;  3U1K A;  5L3J A;  4DR1 I;  3CDI A;  5DRP A;  1Z0E A;  4UTG A;  2O3Z A;  3PS2 A;  2JEA B;  2J7K A;  4YY3 E;  3ZZT A;  1Z5C A;  2C37 B;  1N33 I;  4NXM E;  4DV5 I;  4LFB I;  1IBM E;  4MDT A;  1Z0T A;  2X36 A;  4MU3 A;  2Z5C A;  4USM A;  4OX9 E;  3ZZ0 A;  4HAC A;  4YY3 I;  2PO2 B;  2BA0 D;  3LPS A;  3GLL A;  1XNQ I;  2UU9 E;  3T1H E;  5FLX Q;  4B3S I;  1HNZ E;  2V2V A;  1J5E E;  4IZJ B;  2LJP A;  1RHY A;  1N0U A;  2UXB I;  3GME A;  2ZIT A;  4B3T E;  4LF7 I;  3K1J A;  1Z0G A;  1OJ4 A;  5C0X F;  1FI4 A;  5K36 D;  1ELO A;  5LMN I;  2BM0 A;  1N32 I;  4UT4 A;  3OTO E;  2AE8 A;  4DU8 A;  4FW3 A;  1R6L A;  3ZM7 A;  4MYT A;  2C39 B;  2V2Q A;  4JI4 I;  3HKM A;  1Z0B A;  2C38 A;  2PNZ A;  3UHM A;  1R6M A;  5GMD A;  5JEA C;  1Z5A A;  1USV A;  4LFA I;  5EKW A;  3WU4 A;  2E1R A;  3P06 A;  1OYR A;  4JYA I;  2BA0 G;  3NZK A;  1HNW E;  2UXD I;  4AQY E;  4DUZ E;  1HNW I;  2R3V A;  2Z5E A;  1ZM9 A;  4DR7 I;  2ZM6 I;  4YHH I;  4FW5 A;  4K0K E;  4B3M E;  4MU1 A;  3B82 A;  5C0X B;  2J65 A;  2JEB A;  2DEJ A;  1MX0 A;  3J7A G;  3HJC A;  2F4V I;  4GKJ I;  4JI4 E;  2VQF E;  2JE6 B;  2NN6 A;  2UUC I;  1Z59 A;  4LCH A;  1NHH A;  3PYG A;  1UDS A;  2A2D A;  1ZM3 A;  5LMU E;  3M7N D;  3PRY A;  4JI7 E;  4JV5 E;  4DV1 E;  2GO3 A;  4P7A A;  1USU A;  1XMQ I;  2UXB E;  2NPF A;  2EFG A;  4NXN E;  2CG9 A;  4OX9 I;  2UXC I;  5IWA I;  1J5E I;  1KVK A;  1ZM2 A;  2BR2 B;  1FWK A;  4JI8 E;  4LFB E;  1XNQ E;  5K36 C;  3H4L A;  2HFS A;  1UDN A;  5K36 A;  1XNR I;  2A2C A;  3M6A A;  4DR2 I;  4FW7 A;  2PNL A;  4DR6 E;  2PO0 B;  4GQU A;  2C39 A;  4IZJ D;  1FNM A;  3LTO A;  2R42 A;  1N0V C;  1UEK A;  1VIS A;  1PIE A;  5DNX A;  4DV0 I;  2PO1 B;  1KIJ A;  4OO1 B;  3Q6M A;  2CZ9 A;  4J3D A;  3P76 A;  4DUY I;  4IZK A;  4I3H A;  1XMQ E;  4X65 I;  2HKE A;  1FJG E;  4JI1 E;  1FWL A;  1KKH A;  5C0X E;  3L7Z B;  4JI3 I;  4JI5 E;  5C0X D;  3P3G A;  1IBL E;  2C37 A;  4OZE A;  2BV3 A;  1NHI A;  4DPX A;  4HYM A;  4GKK E;  1HNX I;  1P42 A;  4R1F A;  3V5R A;  1XHK A;  3B4T A;  4B3M I;  4QNK A;  4X66 I;  4JI2 E;  2JEB B;  2PNM A;  2NN6 F;  3T1Y I;  1FKA E;  2O1V A;  3PS1 A;  4IZJ A;  1NZ0 A;  2NN6 D;  3B78 A;  4FW4 A;  3KRN A;  4AQY I;  4DPY A;  3ZKD A;  3J80 C;  4GKK I;  4IFD C;  4JI0 E;  4RPF A;  3QT6 A;  3K17 A;  3WU3 A;  2IER A;  3QT8 A;  1XMO I;  4LFC E;  3J81 C;  2CVE A;  4KQV A;  1Y5Y A;  4DV6 I;  1IBK I;  4B3R I;  4JI5 I;  1EA6 A;  4FW6 A;  3M85 G;  1ZXM A;  5K36 E;  1N32 E;  1H74 A;  2JEA A;  1N34 I;  2E5L E;  2UUA I;  4LF9 E;  4PRV A;  4B3T I;  4LF7 E;  4ISA A;  2VQE E;  4FWH A;  3CWV A;  5K36 B;  4QNJ A;  4URL A;  4X64 I;  3J7A M;  4BA1 B;  4LF4 E;  2GQ0 A;  2IOQ A;  4DPT A;  4DV2 E;  2HKJ A;  4MYU A;  5JEA B;  4X65 E;  4B3S E;  4DR4 I;  3J81 Q;  2O1T A;  4ZVI A;  4DUY E;  1KTV A;  2NN6 C;  5GME A;  4DR3 E;  4DV0 E;  1VI7 A;  4JG4 A;  1XNR E;  5DRQ A;  1A6F A;  2WP8 B;  2CGE A;  4DV7 E;  1HK7 A;  3P3C A;  2UU9 I;  4DR2 E;  4DR6 I;  2Z5B B;  3V2U C;  3Q6N A;  2WNR A;  4MU4 A;  3GON A;  3ZZU A;  2OI2 A;  4NXM I;  1I94 E;  4DR3 I;  4DV4 I;  1N33 E;  4KHP I;  3JAM Q;  2JE6 A;  5C0X C;  5DRR A;  1OYP A;  4DR4 E;  4DV7 I;  1Y4U A;  4DV2 I;  1HR0 E;  4OO1 D;  4Z7C A;  4DV1 I;  1H73 A;  4WUB A;  4X64 E; 
#chains in the Genus database with same CATH homology
 2J65 A;  4J3D A;  4FW6 A;  5DRQ A;  5DRP A;  3P3E A;  3U1Y A;  3P3C A;  2O3Z A;  3PS2 A;  3PS1 A;  4LCH A;  5DRO A;  3NZK A;  4ISA A;  2GO4 A;  4MDT A;  1XXE A;  1YHC A;  4FW4 A;  4U3B A;  3PS3 A;  2IES A;  4U3D A;  4MQY A;  4FW3 A;  4OKG A;  4FW7 A;  3P3G A;  1YH8 A;  2GO3 A;  4IS9 A;  4LCF A;  5DRR A;  4OZE A;  3UHM A;  2IER A;  4FW5 A;  2VES A;  4LCG A;  1P42 A;  2JT2 A;  3P76 A; 
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similar chains in the Genus database (?% sequence similarity)
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similar chains in the pdb database (?% sequence similarity)

 
#similar chains in the Genus database (?% sequence similarity)
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#similar chains, but unknotted
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#similar chains in the pdb database (?% sequence similarity)
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