2IERA

Crystal structure of aquifex aeolicus lpxc complexed with uridine 5'-diphosphate
Total Genus 89
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The genus trace: a function that shows values of genus (vertical axis) for subchains spanned between the first residue, and all other residues (shown on horizontal axis). The number of the latter residue and the genus of a given subchain are shown interactively.

Total Genus
89
sequence length
267
structure length
267
Chain Sequence
b'GLEKTVKEKLSFEGVGIHTGEYSKLIIHPEKEGTGIRFFKNGVYIPARHEFVVHTNHSTDLGFKGQRIKTVEHILSVLHLLEITNVTIEVIGNEIPILDGSGWEFYEAIRKNILNQNREIDYFVVEEPIIVEDEGRLIKAEPSDTLEVTYEGEFKNFLGRQKFTFVEGNEEEIVLARTFAFDWEIEHIKKVGLGKGGSLKNTLVLGKDKVYNPEGLRYENEPVRHKVFDLIGDLYLLGSPVKGKFYSFRGGHSLNVKLVKELAKKQK'
Loading diagram...

The genus matrix. At position (x,y) a genus value for a subchain spanned between x’th and y’th residue is shown. Values of the genus are represented by color, according to the scale given on the right.

Structure visualization

After clicking on a point (x,y) in the genus matrix above, a subchain from x to y is shown in color.

Chord Diagram
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molecule keywords UDP-3-O-[3-hydroxymyristoyl] N-acetylglucosamine deacetylase
publication title Binding of Uridine 5-Diphosphate in the Basic Patch of the Zinc Metalloenzyme Deacetylase LpxC and Implications for Substrate Binding
pubmed doi rcsb
source organism Aquifex aeolicus
molecule tags Hydrolase
total genus 89
structure length 267
sequence length 267
chains with identical sequence B
ec nomenclature ec 3.5.1.108: UDP-3-O-acyl-N-acetylglucosamine deacetylase.
pdb deposition date 2006-09-19

pfam database annotations

chain Pfam Accession Code Pfam Family Identifier Pfam Description
A PF03331 LpxC UDP-3-O-acyl N-acetylglycosamine deacetylase
Image from the rcsb pdb (www.rcsb.org)
cath code
ClassArchitectureTopologyHomologyDomain
3.30.230.20 Alpha Beta 2-Layer Sandwich Ribosomal Protein S5; domain 2 lpxc deacetylase, domain 1 2ierA01
3.30.1700.10 Alpha Beta 2-Layer Sandwich lpxc deacetylase, domain 1 lpxc deacetylase, domain 2 2ierA02
4LCFA 4FW7A 4FW4A 3PS3A 3P3GA 4FW6A 1P42A 3PS2A 3U1YA 2JT2A 3PS1A 3P3EA 3P76A 5DROA 2IESA 2O3ZA 2VESA 4U3BA 5DRQA 5DRPA 4U3DA 4J3DA 4ISAA 2J65A 4MQYA 2GO4A 4IS9A 4OZEA 4FW5A 4LCGA 2IERA 2GO3A 4FW3A 3UHMA 1YHCA 3P3CA 4LCHA 1YH8A 5DRRA 1XXEA 3NZKA 4MDTA 4OKGA
chains in the Genus database with same CATH superfamily
4LF9E 3Q6NA 1B63A 3ZZ0A 4AQYI 1RHYA 2GQ0A 4UT4A 2HFSA 3P3EA 1IBKI 1VI7A 4DR6E 4IFDC 1Z5CA 4DV6I 3WU4A 4BA1A 4X64E 3J81C 3KRNA 4J3DA 2PO1B 2O1TA 3T1YI 4DR1I 1Z59A 1UDNA 1H72C 5K36D 1B62A 2BR2B 4DV5E 4PU9A 1UDOA 4B3TE 5C0XC 3UHMA 4DR2E 4DPXA 4DPYA 4GKJI 3NZKA 4YY3E 4MDTA 3J81Q 5BR8I 4IFDF 4LFCE 5C0XB 4DR4E 2UU9E 1HNZI 5LL6R 4X62I 2OI2A 5DNXA 1USUA 2CG9A 2Z5EA 3U1KA 4JI7I 4LF6E 4KHPI 4LPFA 4YHHI 4JV5E 1ZM3A 1HR0I 1Y6ZA 2A2CA 4IFDD 2HK2A 4DV4E 2O1UA 4OX9I 1MU5A 2V2ZA 5JEAA 1Z0EA 1KIJA 4DR6I 2CGEA 2UXBE 4B3RI 4JYAI 2JEAA 2R3VA 1BKNA 2UUBI 4HYMA 4AIDA 3J80C 2WW4A 4BA2B 2O1VA 4DR2I 1N32E 4DR7I 2V2QA 2VESA 4LF5I 4LFAE 4X66E 4YY3I 5IWAI 4PRVA 1N33E 4MQYA 3GLLA 2UU9I 5A2QC 4DR4I 2VF3A 1HNWI 1VISA 3ZKBA 4ZVIA 4JI5E 4MU0A 4X65I 3M7NG 2PO2A 5JEAF 3M85G 4DV2I 3QT7A 4DR3E 3T1HE 3PS3A 4JI2E 5FLXQ 3QT5A 2HHHI 1FWKA 2HFUA 2CZ9A 2EFGA 3CDIA 3PYGA 2O3ZA 4U3BA 5DRPA 4HACA 5JEAD 4DV0I 1KKHA 1XNRE 1U2RA 4OO1A 4LF7I 2E1RA 5FWKA 5ELWA 5IT9C 1UDSA 1E3PA 1PKPA 4RPFA 3V2UC 1N32I 1S4EA 2HK3A 1XXEA 2WNRA 5K36A 4OKGA 4I3HA 3B78A 4OO1C 1N36E 2C39B 4XTJA 1XHKA 1RREA 2JT2A 3L7ZA 2XEXA 4JI5I 4LFBI 4GKKI 1Y4UA 1RR9A 1HK7A 3M7ND 1R6MA 3M85D 4DU8A 1ZXNA 2LJPA 3WU3A 2F4VE 2DEJA 1H73A 4P7AA 3T1HI 2C38B 5L3JA 4DUZI 3PYEA 4OO1F 4JI8E 3ZZUA 3QT8A 1N0VC 3LTOA 1Y60A 4HXZA 5LMUE 2PO1A 1DARA 3OTOE 2UUAI 4IFDA 2NN6E 3ZKDA 4NXNE 3V5RA 2BR2A 1EA6A 2HKJA 2ZITA 4LF4E 1Z0BA 4GQUA 3P3GA 4FW6A 5K36E 2NN6B 3PS2A 2CVEA 1UDQA 1N0UA 4KQVA 1Z0GA 1FNMA 1OYSA 4OO1D 5DRQA 5C0XA 1HNXE 1ZM4A 4DV1E 1ZM9A 4JI0I 2Z5BB 2J7KA 2IERA 4B3ME 2PO0B 4DU7A 3HULA 4LF8I 1E3HA 2UUCE 2VQEE 4Z7YA 3Q1QA 2AJ4A 4FN5A 1IBME 4IZJB 4IFDE 1ZM2A 4NXME 5LMNE 2C37B 4FWGA 1NHJA 1S16A 4LF4I 5C0XF 1FJGI 4LF9I 4BA2A 4PRXA 1J5EE 1ZXMA 4NBQA 4DV3E 2PNZB 1XMQE 2ZM6E 4X64I 3CWVA 4DV1I 4DPWA 4AM3A 4USKA 5C0XD 4DV5I 4QNKA 4B3MI 1I94E 4B3TI 1XNQE 2IOQA 3WU6A 2UUCI 4FWDA 1PIEA 4DUYE 3ZM7A 1N34E 2DEIA 4N3OA 2O1WA 4WUDA 4LFCI 1Z5BA 5FLXC 4JI4E 4NXMI 4LCHA 1Z0VA 2R42A 5EL9A 4DPUA 4JI6E 1XMOE 4P52A 4LF6I 2JEBA 5JEAE 4JV5I 2UXCE 2VQFE 3U1YA 1FWLA 2BA1G 4DV4I 4B3SE 3WU5A 1XMQI 4R1FA 2BA0G 2UXBI 4WUBA 2C39A 1H74A 2J65A 2JEAB 4FW5A 2GO3A 2BV3A 1XNQI 4MU1A 1YHCA 2WP8B 2C38A 4LFAI 4X66I 1N34I 2UXDE 1N33I 1Y4SA 1Z0TA 3J7AG 1QZRA 3K1JA 4JI6I 2PO2B 1XMOI 4DR3I 3JAMC 3B4TA 2JE6B 2Z5CB 4AQYE 4WUCA 4OZEA 2F1DA 2VQFI 4DR5E 3PYDA 4LCGA 4JI2I 4B3SI 4USMA 3DD6A 1FI4A 1MX0A 4OO1E 2BA0D 3T1YE 2NN6C 5GMDA 4HY1A 3ZZTA 3JAMQ 4FW4A 1XNRI 3J7AM 1KTVA 2BM0A 5BR8E 1FKAE 5K36B 2UXDI 1HNZE 1ELOA 2C37A 2HKEA 5EKWA 1N36I 3K85A 2NN6F 1Y5YA 4X62E 1Z0WA 3H4LA 2AE8A 3B8HA 2BM1A 4JI7E 4KHPE 4YHHE 3LNUA 1HR0E 4LCFA 2PNMA 2E5LE 2F4VI 4JI3E 4DR5I 4OX9E 2PNZA 3CDJA 1OYPA 2V8PA 3QT6A 3PS1A 1Z0CA 2NN6D 3GCMA 4JI8I 1WUUA 4B3RE 4JYAE 1NHHA 4M1KA 5LMUI 3OTOI 2PNLA 3M6AA 1R6LA 4IZKB 4NXNI 4IFDB 2UUBE 2GO4A 4DV7E 5C0XE 1KVKA 1OJ4A 1H7UA 4UTGA 4DR7E 4LF5E 2A2DA 5IWAE 1HNXI 4BA1B 1EI1A 1HNWE 1P42A 4MYTA 4X65E 4IZJD 2GS8A 4DV2E 4URLA 3P76A 3GMEA 2VQEI 2NPFA 2E5LI 4JI3I 4MU3A 4DPTA 2HHHE 4LOMA 1IBMI 4FW3A 5LMNI 3LPSA 3PYFA 4DV0E 4QNJA 4LF7E 4EMDA 3K17A 4Z7CA 4DV7I 5JEAC 2X7IA 3B82A 2WP8A 3GONA 1J5EI 4DV3I 5DROA 2IESA 1OYRA 5JEAB 2ZM6I 4JI1E 3Q6MA 3J80Q 1NZ0A 2Q2EB 4LFBE 4GKKE 1I94I 4DXLA 3P3CA 4FW9A 2JE6A 2Z5CA 5GMEA 3P06A 4K0KE 4DUYI 2ZBKB 5A2QQ 4DUZE 1UEKA 3PRYA 4JI4I 1IBLE 2UUAE 4MU4A 2V34A 1A6FA 2IOPA 4FWHA 1USVA 4ISAA 1MG7A 2NN6A 1NHIA 4AIMA 2JEBB 2UXCI 1IBKE 2BA1D 4DV6E 1YH8A 5DRRA 3D4JA 4JI1I 2Z5BA 4JI0E 4FW7A 4DR1E 5DNLA 3F0NA 5IT9Q 4HACB 4MYUA 2V2VA 2PO0A 4OO1B 3H1CA 1H7SA 5K36C 4LF8E 4GKJE 4U3DA 3HJCA 4K0KI 2WNRB 4IZJA 4IS9A 1Z5AA 4JG4A 1K47A 4ED4A 1IBLI 4IZKA 3HKMA 1D6TA 2X36A 3L7ZB 1FJGE 5K36F 1PVGA
chains in the Genus database with same CATH topology
4LCFA 4FW7A 4FW4A 3PS3A 3P3GA 4FW6A 1P42A 3PS2A 3U1YA 2JT2A 3PS1A 3P3EA 3P76A 5DROA 2IESA 2O3ZA 2VESA 4U3BA 5DRQA 5DRPA 4U3DA 4J3DA 4ISAA 2J65A 4MQYA 2GO4A 4IS9A 4OZEA 4FW5A 4LCGA 2IERA 2GO3A 4FW3A 3UHMA 1YHCA 3P3CA 4LCHA 1YH8A 5DRRA 1XXEA 3NZKA 4MDTA 4OKGA
chains in the Genus database with same CATH homology


 
#chains in the Genus database with same CATH superfamily
 4LCF A;  4FW7 A;  4FW4 A;  3PS3 A;  3P3G A;  4FW6 A;  1P42 A;  3PS2 A;  3U1Y A;  2JT2 A;  3PS1 A;  3P3E A;  3P76 A;  5DRO A;  2IES A;  2O3Z A;  2VES A;  4U3B A;  5DRQ A;  5DRP A;  4U3D A;  4J3D A;  4ISA A;  2J65 A;  4MQY A;  2GO4 A;  4IS9 A;  4OZE A;  4FW5 A;  4LCG A;  2IER A;  2GO3 A;  4FW3 A;  3UHM A;  1YHC A;  3P3C A;  4LCH A;  1YH8 A;  5DRR A;  1XXE A;  3NZK A;  4MDT A;  4OKG A; 
#chains in the Genus database with same CATH topology
 4LF9 E;  3Q6N A;  1B63 A;  3ZZ0 A;  4AQY I;  1RHY A;  2GQ0 A;  4UT4 A;  2HFS A;  3P3E A;  1IBK I;  1VI7 A;  4DR6 E;  4IFD C;  1Z5C A;  4DV6 I;  3WU4 A;  4BA1 A;  4X64 E;  3J81 C;  3KRN A;  4J3D A;  2PO1 B;  2O1T A;  3T1Y I;  4DR1 I;  1Z59 A;  1UDN A;  1H72 C;  5K36 D;  1B62 A;  2BR2 B;  4DV5 E;  4PU9 A;  1UDO A;  4B3T E;  5C0X C;  3UHM A;  4DR2 E;  4DPX A;  4DPY A;  4GKJ I;  3NZK A;  4YY3 E;  4MDT A;  3J81 Q;  5BR8 I;  4IFD F;  4LFC E;  5C0X B;  4DR4 E;  2UU9 E;  1HNZ I;  5LL6 R;  4X62 I;  2OI2 A;  5DNX A;  1USU A;  2CG9 A;  2Z5E A;  3U1K A;  4JI7 I;  4LF6 E;  4KHP I;  4LPF A;  4YHH I;  4JV5 E;  1ZM3 A;  1HR0 I;  1Y6Z A;  2A2C A;  4IFD D;  2HK2 A;  4DV4 E;  2O1U A;  4OX9 I;  1MU5 A;  2V2Z A;  5JEA A;  1Z0E A;  1KIJ A;  4DR6 I;  2CGE A;  2UXB E;  4B3R I;  4JYA I;  2JEA A;  2R3V A;  1BKN A;  2UUB I;  4HYM A;  4AID A;  3J80 C;  2WW4 A;  4BA2 B;  2O1V A;  4DR2 I;  1N32 E;  4DR7 I;  2V2Q A;  2VES A;  4LF5 I;  4LFA E;  4X66 E;  4YY3 I;  5IWA I;  4PRV A;  1N33 E;  4MQY A;  3GLL A;  2UU9 I;  5A2Q C;  4DR4 I;  2VF3 A;  1HNW I;  1VIS A;  3ZKB A;  4ZVI A;  4JI5 E;  4MU0 A;  4X65 I;  3M7N G;  2PO2 A;  5JEA F;  3M85 G;  4DV2 I;  3QT7 A;  4DR3 E;  3T1H E;  3PS3 A;  4JI2 E;  5FLX Q;  3QT5 A;  2HHH I;  1FWK A;  2HFU A;  2CZ9 A;  2EFG A;  3CDI A;  3PYG A;  2O3Z A;  4U3B A;  5DRP A;  4HAC A;  5JEA D;  4DV0 I;  1KKH A;  1XNR E;  1U2R A;  4OO1 A;  4LF7 I;  2E1R A;  5FWK A;  5ELW A;  5IT9 C;  1UDS A;  1E3P A;  1PKP A;  4RPF A;  3V2U C;  1N32 I;  1S4E A;  2HK3 A;  1XXE A;  2WNR A;  5K36 A;  4OKG A;  4I3H A;  3B78 A;  4OO1 C;  1N36 E;  2C39 B;  4XTJ A;  1XHK A;  1RRE A;  2JT2 A;  3L7Z A;  2XEX A;  4JI5 I;  4LFB I;  4GKK I;  1Y4U A;  1RR9 A;  1HK7 A;  3M7N D;  1R6M A;  3M85 D;  4DU8 A;  1ZXN A;  2LJP A;  3WU3 A;  2F4V E;  2DEJ A;  1H73 A;  4P7A A;  3T1H I;  2C38 B;  5L3J A;  4DUZ I;  3PYE A;  4OO1 F;  4JI8 E;  3ZZU A;  3QT8 A;  1N0V C;  3LTO A;  1Y60 A;  4HXZ A;  5LMU E;  2PO1 A;  1DAR A;  3OTO E;  2UUA I;  4IFD A;  2NN6 E;  3ZKD A;  4NXN E;  3V5R A;  2BR2 A;  1EA6 A;  2HKJ A;  2ZIT A;  4LF4 E;  1Z0B A;  4GQU A;  3P3G A;  4FW6 A;  5K36 E;  2NN6 B;  3PS2 A;  2CVE A;  1UDQ A;  1N0U A;  4KQV A;  1Z0G A;  1FNM A;  1OYS A;  4OO1 D;  5DRQ A;  5C0X A;  1HNX E;  1ZM4 A;  4DV1 E;  1ZM9 A;  4JI0 I;  2Z5B B;  2J7K A;  2IER A;  4B3M E;  2PO0 B;  4DU7 A;  3HUL A;  4LF8 I;  1E3H A;  2UUC E;  2VQE E;  4Z7Y A;  3Q1Q A;  2AJ4 A;  4FN5 A;  1IBM E;  4IZJ B;  4IFD E;  1ZM2 A;  4NXM E;  5LMN E;  2C37 B;  4FWG A;  1NHJ A;  1S16 A;  4LF4 I;  5C0X F;  1FJG I;  4LF9 I;  4BA2 A;  4PRX A;  1J5E E;  1ZXM A;  4NBQ A;  4DV3 E;  2PNZ B;  1XMQ E;  2ZM6 E;  4X64 I;  3CWV A;  4DV1 I;  4DPW A;  4AM3 A;  4USK A;  5C0X D;  4DV5 I;  4QNK A;  4B3M I;  1I94 E;  4B3T I;  1XNQ E;  2IOQ A;  3WU6 A;  2UUC I;  4FWD A;  1PIE A;  4DUY E;  3ZM7 A;  1N34 E;  2DEI A;  4N3O A;  2O1W A;  4WUD A;  4LFC I;  1Z5B A;  5FLX C;  4JI4 E;  4NXM I;  4LCH A;  1Z0V A;  2R42 A;  5EL9 A;  4DPU A;  4JI6 E;  1XMO E;  4P52 A;  4LF6 I;  2JEB A;  5JEA E;  4JV5 I;  2UXC E;  2VQF E;  3U1Y A;  1FWL A;  2BA1 G;  4DV4 I;  4B3S E;  3WU5 A;  1XMQ I;  4R1F A;  2BA0 G;  2UXB I;  4WUB A;  2C39 A;  1H74 A;  2J65 A;  2JEA B;  4FW5 A;  2GO3 A;  2BV3 A;  1XNQ I;  4MU1 A;  1YHC A;  2WP8 B;  2C38 A;  4LFA I;  4X66 I;  1N34 I;  2UXD E;  1N33 I;  1Y4S A;  1Z0T A;  3J7A G;  1QZR A;  3K1J A;  4JI6 I;  2PO2 B;  1XMO I;  4DR3 I;  3JAM C;  3B4T A;  2JE6 B;  2Z5C B;  4AQY E;  4WUC A;  4OZE A;  2F1D A;  2VQF I;  4DR5 E;  3PYD A;  4LCG A;  4JI2 I;  4B3S I;  4USM A;  3DD6 A;  1FI4 A;  1MX0 A;  4OO1 E;  2BA0 D;  3T1Y E;  2NN6 C;  5GMD A;  4HY1 A;  3ZZT A;  3JAM Q;  4FW4 A;  1XNR I;  3J7A M;  1KTV A;  2BM0 A;  5BR8 E;  1FKA E;  5K36 B;  2UXD I;  1HNZ E;  1ELO A;  2C37 A;  2HKE A;  5EKW A;  1N36 I;  3K85 A;  2NN6 F;  1Y5Y A;  4X62 E;  1Z0W A;  3H4L A;  2AE8 A;  3B8H A;  2BM1 A;  4JI7 E;  4KHP E;  4YHH E;  3LNU A;  1HR0 E;  4LCF A;  2PNM A;  2E5L E;  2F4V I;  4JI3 E;  4DR5 I;  4OX9 E;  2PNZ A;  3CDJ A;  1OYP A;  2V8P A;  3QT6 A;  3PS1 A;  1Z0C A;  2NN6 D;  3GCM A;  4JI8 I;  1WUU A;  4B3R E;  4JYA E;  1NHH A;  4M1K A;  5LMU I;  3OTO I;  2PNL A;  3M6A A;  1R6L A;  4IZK B;  4NXN I;  4IFD B;  2UUB E;  2GO4 A;  4DV7 E;  5C0X E;  1KVK A;  1OJ4 A;  1H7U A;  4UTG A;  4DR7 E;  4LF5 E;  2A2D A;  5IWA E;  1HNX I;  4BA1 B;  1EI1 A;  1HNW E;  1P42 A;  4MYT A;  4X65 E;  4IZJ D;  2GS8 A;  4DV2 E;  4URL A;  3P76 A;  3GME A;  2VQE I;  2NPF A;  2E5L I;  4JI3 I;  4MU3 A;  4DPT A;  2HHH E;  4LOM A;  1IBM I;  4FW3 A;  5LMN I;  3LPS A;  3PYF A;  4DV0 E;  4QNJ A;  4LF7 E;  4EMD A;  3K17 A;  4Z7C A;  4DV7 I;  5JEA C;  2X7I A;  3B82 A;  2WP8 A;  3GON A;  1J5E I;  4DV3 I;  5DRO A;  2IES A;  1OYR A;  5JEA B;  2ZM6 I;  4JI1 E;  3Q6M A;  3J80 Q;  1NZ0 A;  2Q2E B;  4LFB E;  4GKK E;  1I94 I;  4DXL A;  3P3C A;  4FW9 A;  2JE6 A;  2Z5C A;  5GME A;  3P06 A;  4K0K E;  4DUY I;  2ZBK B;  5A2Q Q;  4DUZ E;  1UEK A;  3PRY A;  4JI4 I;  1IBL E;  2UUA E;  4MU4 A;  2V34 A;  1A6F A;  2IOP A;  4FWH A;  1USV A;  4ISA A;  1MG7 A;  2NN6 A;  1NHI A;  4AIM A;  2JEB B;  2UXC I;  1IBK E;  2BA1 D;  4DV6 E;  1YH8 A;  5DRR A;  3D4J A;  4JI1 I;  2Z5B A;  4JI0 E;  4FW7 A;  4DR1 E;  5DNL A;  3F0N A;  5IT9 Q;  4HAC B;  4MYU A;  2V2V A;  2PO0 A;  4OO1 B;  3H1C A;  1H7S A;  5K36 C;  4LF8 E;  4GKJ E;  4U3D A;  3HJC A;  4K0K I;  2WNR B;  4IZJ A;  4IS9 A;  1Z5A A;  4JG4 A;  1K47 A;  4ED4 A;  1IBL I;  4IZK A;  3HKM A;  1D6T A;  2X36 A;  3L7Z B;  1FJG E;  5K36 F;  1PVG A; 
#chains in the Genus database with same CATH homology
 4LCF A;  4FW7 A;  4FW4 A;  3PS3 A;  3P3G A;  4FW6 A;  1P42 A;  3PS2 A;  3U1Y A;  2JT2 A;  3PS1 A;  3P3E A;  3P76 A;  5DRO A;  2IES A;  2O3Z A;  2VES A;  4U3B A;  5DRQ A;  5DRP A;  4U3D A;  4J3D A;  4ISA A;  2J65 A;  4MQY A;  2GO4 A;  4IS9 A;  4OZE A;  4FW5 A;  4LCG A;  2IER A;  2GO3 A;  4FW3 A;  3UHM A;  1YHC A;  3P3C A;  4LCH A;  1YH8 A;  5DRR A;  1XXE A;  3NZK A;  4MDT A;  4OKG A; 
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similar chains in the Genus database (?% sequence similarity)
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similar chains in the pdb database (?% sequence similarity)

 
#similar chains in the Genus database (?% sequence similarity)
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#similar chains, but unknotted
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#similar chains in the pdb database (?% sequence similarity)
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