2VESA

Crystal structure of lpxc from pseudomonas aeruginosa complexed with the potent bb-78485 inhibitor
Total Genus 88
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The genus trace: a function that shows values of genus (vertical axis) for subchains spanned between the first residue, and all other residues (shown on horizontal axis). The number of the latter residue and the genus of a given subchain are shown interactively.

Total Genus
88
sequence length
294
structure length
294
Chain Sequence
b'MIKQRTLKNIIRATGVGLHSGEKVYLTLKPAPVDTGIVFSRTDLDPVVEIPARAENVGETTMSTTLVKGDVKVDTVEHLLSAMAGLGIDNAYVELSASEVPIMDGSAGPFVFLIQSAGLQEQEAAKKFIRIKREVSVEEGDKRAVFVPFDGFKVSFEIDFDHPVFRGRTQQASVDFSSTSFVKEVSRARTFGFMRDIEYLRSQNLALGGSVENAIVVDENRVLNEDGLRYEDEFVKHKILDAIGDLYLLGNSLIGEFRGFKSGHALNNQLLRTLIADKDAWEVVTFEDARTAPI'
Loading diagram...

The genus matrix. At position (x,y) a genus value for a subchain spanned between x’th and y’th residue is shown. Values of the genus are represented by color, according to the scale given on the right.

Structure visualization

After clicking on a point (x,y) in the genus matrix above, a subchain from x to y is shown in color.

Chord Diagram
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molecule keywords UDP-3-O-acyl-N-acetylglucosamine deacetylase
publication title Crystal structure of LpxC from Pseudomonas aeruginosa complexed with the potent BB-78485 inhibitor.
pubmed doi rcsb
source organism Pseudomonas aeruginosa
molecule tags Hydrolase
total genus 88
structure length 294
sequence length 294
chains with identical sequence B, C
ec nomenclature ec 3.5.1.108: UDP-3-O-acyl-N-acetylglucosamine deacetylase.
pdb deposition date 2007-10-26

pfam database annotations

chain Pfam Accession Code Pfam Family Identifier Pfam Description
A PF03331 LpxC UDP-3-O-acyl N-acetylglycosamine deacetylase
Image from the rcsb pdb (www.rcsb.org)
cath code
ClassArchitectureTopologyHomologyDomain
3.30.230.20 Alpha Beta 2-Layer Sandwich Ribosomal Protein S5; domain 2 lpxc deacetylase, domain 1 2vesA01
3.30.1700.10 Alpha Beta 2-Layer Sandwich lpxc deacetylase, domain 1 lpxc deacetylase, domain 2 2vesA02
3P3GA 4ISAA 4LCGA 4LCHA 2J65A 2VESA 4MQYA 4MDTA 4U3DA 4IS9A 2GO4A 3P3EA 4U3BA 4FW3A 4FW5A 1YHCA 1YH8A 4FW6A 3PS1A 3P76A 1P42A 3PS3A 2IERA 1XXEA 5DRRA 3NZKA 4FW7A 4OKGA 4OZEA 3U1YA 3PS2A 2JT2A 3UHMA 4LCFA 4FW4A 2O3ZA 5DRPA 2IESA 4J3DA 2GO3A 5DRQA 3P3CA 5DROA
chains in the Genus database with same CATH superfamily
4ISAA 2OI2A 4OO1E 4FWHA 3ZZUA 4JI0I 4ZVIA 4IS9A 5IWAI 1XMOE 4LF8E 1UEKA 2GO4A 4DR4E 1XNQI 4KHPE 4DV7E 3HULA 1PVGA 2BR2B 1E3PA 1OYPA 4JG4A 1N32I 3CDJA 3QT7A 4JV5E 4GKKE 5IT9C 5C0XF 2E5LI 1N0UA 1S16A 4XTJA 2PO0B 1NHHA 1UDSA 4FW4A 2XEXA 2O1UA 4JYAE 5JEAF 1Z0EA 4LCHA 4DR1E 2C38A 5LMUI 4LF5I 4WUDA 3CDIA 1Z0CA 1QZRA 3J7AG 4DR3E 4JI4I 4DR2I 1VISA 4JI6E 4FW3A 4IFDF 4JI1I 4DV6I 2V2VA 2X7IA 3J80C 2UXDE 1ZM4A 1OYSA 4P52A 4HACA 3T1YI 4DR7E 5K36F 4BA1A 4LF6E 3P06A 1B63A 2JT2A 3H1CA 2R42A 3Q1QA 3ZZ0A 2C37B 1HNXE 2WP8A 3KRNA 2ZM6E 1VI7A 4X66E 1J5EI 4LF4E 2A2DA 1MX0A 1HNWI 4NXNE 4B3RI 3K17A 4GKJI 1XNRE 1YHCA 1FJGE 4DV2E 4X62E 1ZXMA 1EI1A 4DV0I 1XMQI 5C0XA 4OKGA 1USVA 4R1FA 4PRXA 1XMOI 4LF8I 1H74A 4DR4I 2C38B 4KHPI 4DV7I 4HY1A 3WU4A 2BV3A 2PNZA 1K47A 2PO2B 1R6MA 3M85D 4JV5I 3M85G 4GKKI 5JEAA 2ZITA 4JI2E 5LMNE 1ZM3A 2GS8A 4M1KA 4B3SI 1Y4UA 5JEAD 2Z5CB 4HACB 4FW5A 3JAMQ 4FW6A 2BA0D 4DV5E 4BA2B 2O1WA 4JYAI 4NBQA 2LJPA 4DR1I 3M7ND 2E1RA 4LPFA 1N36I 5C0XC 4DR3I 4UT4A 4JI6I 3PYGA 4YHHI 2HK3A 1IBKI 3T1HE 1RREA 1NHIA 2UXDI 4DR7I 2UXCE 1NZ0A 4LF6I 3ZKBA 5C0XB 2C39B 1N34E 1ZM2A 3OTOE 1HNXI 2ZM6I 4X66I 4IFDC 4LF4I 4OZEA 3ZZTA 4NXNI 4Z7CA 4OO1F 5BR8E 5K36C 1Z0GA 5JEAB 1XNRI 1OJ4A 4AIDA 2BA1D 1ZM9A 1FJGI 2IESA 4DV2I 4X62I 3P3CA 1IBLE 5DROA 3P3GA 1XMQE 1Y4SA 2F4VE 2WNRB 4USKA 4IFDB 1Z5BA 2X36A 2DEJA 4QNKA 4MU0A 1EA6A 5A2QC 1IBME 2V2ZA 5K36B 1WUUA 3QT8A 4AIMA 2IERA 2PO1A 4MU3A 4MYUA 1FWLA 3F0NA 4JI2I 5LMNI 2NN6D 3GCMA 4K0KE 5FLXQ 4URLA 4IZKB 4B3SE 2AJ4A 3B8HA 4QNJA 2HHHE 3LNUA 4DV5I 1ZXNA 1RHYA 1N36E 2Z5BA 2UUBE 2CG9A 4IZJD 2V34A 1Z0TA 1B62A 4DPTA 4YHHE 1IBKE 2JEBA 4DPUA 3T1HI 1H72C 2UXCI 4DV1I 4OO1A 4DU8A 4LCFA 5C0XE 1N34I 4RPFA 4MU1A 3OTOI 4OO1D 4MQYA 4DPXA 4PRVA 5BR8I 4LFBE 2WW4A 2JEAA 3L7ZA 3U1KA 2UU9E 1IBLI 3PYFA 3B82A 1Y6ZA 4N3OA 4B3ME 2Z5EA 2F4VI 4IFDE 4NXME 2HFUA 1HNZE 3J7AM 4DPYA 4MYTA 2O3ZA 2HKJA 1IBMI 5K36E 2UUCE 1BKNA 4DV3E 4LCGA 4DUZE 2VF3A 1Z0VA 4DR5E 4K0KI 3P3EA 3K1JA 2VQEE 4AQYI 2HHHI 4BA1B 4AM3A 2NN6F 5DNLA 1RR9A 4FW7A 3PYDA 3J81Q 1HR0E 2WP8B 1Y5YA 2UUBI 4X65E 1H7SA 4B3TE 1HK7A 4FWDA 2JEAB 3L7ZB 3JAMC 5JEAC 4DV1E 4LFCE 4PU9A 4X64I 2J7KA 2HK2A 3PS1A 2UUAE 1UDQA 1P42A 2DEIA 3LTOA 3QT5A 5DRRA 4LFAE 3GLLA 1N0VC 1N33E 3V2UC 1NHJA 2PNZB 4OX9I 4LFBI 4Z7YA 1PIEA 5DRQA 4I3HA 2UU9I 1ELOA 2ZBKB 4DPWA 2IOPA 4B3MI 1USUA 4U3DA 4NXMI 1HNZI 2VQFE 3HJCA 2NPFA 2JE6A 2UUCI 2NN6A 3GONA 4DV3I 2CGEA 4DUZI 5IT9Q 2CVEA 4DR5I 5DNXA 1MU5A 4JI7E 2EFGA 5EKWA 2VQEI 3LPSA 4DU7A 4AQYE 2J65A 5L3JA 2C37A 2F1DA 4IZJA 4MU4A 4DV4E 1HR0I 4ED4A 1OYRA 4X65I 3K85A 3Q6NA 4YY3E 1FKAE 4UTGA 1H7UA 1KVKA 4FN5A 4B3TI 1PKPA 1XXEA 5FLXC 2BM1A 2NN6C 3QT6A 3J80Q 3PS2A 1A6FA 4LF7E 4DXLA 4JI8E 4LFCI 1DARA 1R6LA 4FW9A 3GMEA 4X64E 2PNMA 4EMDA 2UUAI 1KIJA 4LFAI 2PO2A 2PNLA 1KTVA 4FWGA 2JE6B 5ELWA 4LF9E 2NN6B 1N33I 4JI5E 2V8PA 4U3BA 4OX9E 2PO1B 5JEAE 2UXBE 2O1TA 2GQ0A 3M6AA 2Z5CA 3CWVA 4BA2A 4DR6E 3D4JA 3UHMA 1Z5AA 1I94E 3WU3A 4IZJB 2HFSA 2VQFI 1FI4A 4J3DA 2Z5BB 4OO1C 4LOMA 1FWKA 2VESA 3B78A 4WUCA 1D6TA 4DUYE 1H73A 2JEBB 3ZM7A 4JI3I 5FWKA 4JI7I 1UDOA 1Y60A 4GQUA 2C39A 1UDNA 2Q2EB 3NZKA 4OO1B 1KKHA 4JI0E 5LL6R 3B4TA 3PRYA 1FNMA 5IWAE 4DV4I 1Z5CA 3ZKDA 5C0XD 1XNQE 5EL9A 4USMA 2AE8A 1N32E 4YY3I 2E5LE 1Z0WA 5GMDA 3DD6A 2HKEA 4LF7I 2WNRA 4JI8I 3V5RA 4IFDA 2BA0G 2A2CA 2IOQA 4KQVA 3J81C 3PS3A 5LMUE 2CZ9A 3PYEA 4LF5E 5K36A 3M7NG 4IFDD 1E3HA 2BR2A 1S4EA 4HXZA 1MG7A 3U1YA 4JI4E 2V2QA 3HKMA 4DR2E 2NN6E 2BM0A 3H4LA 4LF9I 4JI5I 5K36D 3WU6A 4JI1E 4DV6E 2PO0A 2UXBI 4IZKA 3T1YE 4P7AA 4MDTA 4DR6I 2R3VA 1I94I 1XHKA 1YH8A 1U2RA 4WUBA 4HYMA 3P76A 5A2QQ 3Q6MA 1J5EE 1Z59A 1HNWE 2O1VA 4B3RE 5GMEA 3WU5A 4DUYI 4GKJE 1Z0BA 4JI3E 2BA1G 5DRPA 2GO3A 4DV0E
chains in the Genus database with same CATH topology
3P3GA 4ISAA 4LCGA 4LCHA 2J65A 2VESA 4MQYA 4MDTA 4U3DA 4IS9A 2GO4A 3P3EA 4U3BA 4FW3A 4FW5A 1YHCA 1YH8A 4FW6A 3PS1A 3P76A 1P42A 3PS3A 2IERA 1XXEA 5DRRA 3NZKA 4FW7A 4OKGA 4OZEA 3U1YA 3PS2A 2JT2A 3UHMA 4LCFA 4FW4A 2O3ZA 5DRPA 2IESA 4J3DA 2GO3A 5DRQA 3P3CA 5DROA
chains in the Genus database with same CATH homology


 
#chains in the Genus database with same CATH superfamily
 3P3G A;  4ISA A;  4LCG A;  4LCH A;  2J65 A;  2VES A;  4MQY A;  4MDT A;  4U3D A;  4IS9 A;  2GO4 A;  3P3E A;  4U3B A;  4FW3 A;  4FW5 A;  1YHC A;  1YH8 A;  4FW6 A;  3PS1 A;  3P76 A;  1P42 A;  3PS3 A;  2IER A;  1XXE A;  5DRR A;  3NZK A;  4FW7 A;  4OKG A;  4OZE A;  3U1Y A;  3PS2 A;  2JT2 A;  3UHM A;  4LCF A;  4FW4 A;  2O3Z A;  5DRP A;  2IES A;  4J3D A;  2GO3 A;  5DRQ A;  3P3C A;  5DRO A; 
#chains in the Genus database with same CATH topology
 4ISA A;  2OI2 A;  4OO1 E;  4FWH A;  3ZZU A;  4JI0 I;  4ZVI A;  4IS9 A;  5IWA I;  1XMO E;  4LF8 E;  1UEK A;  2GO4 A;  4DR4 E;  1XNQ I;  4KHP E;  4DV7 E;  3HUL A;  1PVG A;  2BR2 B;  1E3P A;  1OYP A;  4JG4 A;  1N32 I;  3CDJ A;  3QT7 A;  4JV5 E;  4GKK E;  5IT9 C;  5C0X F;  2E5L I;  1N0U A;  1S16 A;  4XTJ A;  2PO0 B;  1NHH A;  1UDS A;  4FW4 A;  2XEX A;  2O1U A;  4JYA E;  5JEA F;  1Z0E A;  4LCH A;  4DR1 E;  2C38 A;  5LMU I;  4LF5 I;  4WUD A;  3CDI A;  1Z0C A;  1QZR A;  3J7A G;  4DR3 E;  4JI4 I;  4DR2 I;  1VIS A;  4JI6 E;  4FW3 A;  4IFD F;  4JI1 I;  4DV6 I;  2V2V A;  2X7I A;  3J80 C;  2UXD E;  1ZM4 A;  1OYS A;  4P52 A;  4HAC A;  3T1Y I;  4DR7 E;  5K36 F;  4BA1 A;  4LF6 E;  3P06 A;  1B63 A;  2JT2 A;  3H1C A;  2R42 A;  3Q1Q A;  3ZZ0 A;  2C37 B;  1HNX E;  2WP8 A;  3KRN A;  2ZM6 E;  1VI7 A;  4X66 E;  1J5E I;  4LF4 E;  2A2D A;  1MX0 A;  1HNW I;  4NXN E;  4B3R I;  3K17 A;  4GKJ I;  1XNR E;  1YHC A;  1FJG E;  4DV2 E;  4X62 E;  1ZXM A;  1EI1 A;  4DV0 I;  1XMQ I;  5C0X A;  4OKG A;  1USV A;  4R1F A;  4PRX A;  1XMO I;  4LF8 I;  1H74 A;  4DR4 I;  2C38 B;  4KHP I;  4DV7 I;  4HY1 A;  3WU4 A;  2BV3 A;  2PNZ A;  1K47 A;  2PO2 B;  1R6M A;  3M85 D;  4JV5 I;  3M85 G;  4GKK I;  5JEA A;  2ZIT A;  4JI2 E;  5LMN E;  1ZM3 A;  2GS8 A;  4M1K A;  4B3S I;  1Y4U A;  5JEA D;  2Z5C B;  4HAC B;  4FW5 A;  3JAM Q;  4FW6 A;  2BA0 D;  4DV5 E;  4BA2 B;  2O1W A;  4JYA I;  4NBQ A;  2LJP A;  4DR1 I;  3M7N D;  2E1R A;  4LPF A;  1N36 I;  5C0X C;  4DR3 I;  4UT4 A;  4JI6 I;  3PYG A;  4YHH I;  2HK3 A;  1IBK I;  3T1H E;  1RRE A;  1NHI A;  2UXD I;  4DR7 I;  2UXC E;  1NZ0 A;  4LF6 I;  3ZKB A;  5C0X B;  2C39 B;  1N34 E;  1ZM2 A;  3OTO E;  1HNX I;  2ZM6 I;  4X66 I;  4IFD C;  4LF4 I;  4OZE A;  3ZZT A;  4NXN I;  4Z7C A;  4OO1 F;  5BR8 E;  5K36 C;  1Z0G A;  5JEA B;  1XNR I;  1OJ4 A;  4AID A;  2BA1 D;  1ZM9 A;  1FJG I;  2IES A;  4DV2 I;  4X62 I;  3P3C A;  1IBL E;  5DRO A;  3P3G A;  1XMQ E;  1Y4S A;  2F4V E;  2WNR B;  4USK A;  4IFD B;  1Z5B A;  2X36 A;  2DEJ A;  4QNK A;  4MU0 A;  1EA6 A;  5A2Q C;  1IBM E;  2V2Z A;  5K36 B;  1WUU A;  3QT8 A;  4AIM A;  2IER A;  2PO1 A;  4MU3 A;  4MYU A;  1FWL A;  3F0N A;  4JI2 I;  5LMN I;  2NN6 D;  3GCM A;  4K0K E;  5FLX Q;  4URL A;  4IZK B;  4B3S E;  2AJ4 A;  3B8H A;  4QNJ A;  2HHH E;  3LNU A;  4DV5 I;  1ZXN A;  1RHY A;  1N36 E;  2Z5B A;  2UUB E;  2CG9 A;  4IZJ D;  2V34 A;  1Z0T A;  1B62 A;  4DPT A;  4YHH E;  1IBK E;  2JEB A;  4DPU A;  3T1H I;  1H72 C;  2UXC I;  4DV1 I;  4OO1 A;  4DU8 A;  4LCF A;  5C0X E;  1N34 I;  4RPF A;  4MU1 A;  3OTO I;  4OO1 D;  4MQY A;  4DPX A;  4PRV A;  5BR8 I;  4LFB E;  2WW4 A;  2JEA A;  3L7Z A;  3U1K A;  2UU9 E;  1IBL I;  3PYF A;  3B82 A;  1Y6Z A;  4N3O A;  4B3M E;  2Z5E A;  2F4V I;  4IFD E;  4NXM E;  2HFU A;  1HNZ E;  3J7A M;  4DPY A;  4MYT A;  2O3Z A;  2HKJ A;  1IBM I;  5K36 E;  2UUC E;  1BKN A;  4DV3 E;  4LCG A;  4DUZ E;  2VF3 A;  1Z0V A;  4DR5 E;  4K0K I;  3P3E A;  3K1J A;  2VQE E;  4AQY I;  2HHH I;  4BA1 B;  4AM3 A;  2NN6 F;  5DNL A;  1RR9 A;  4FW7 A;  3PYD A;  3J81 Q;  1HR0 E;  2WP8 B;  1Y5Y A;  2UUB I;  4X65 E;  1H7S A;  4B3T E;  1HK7 A;  4FWD A;  2JEA B;  3L7Z B;  3JAM C;  5JEA C;  4DV1 E;  4LFC E;  4PU9 A;  4X64 I;  2J7K A;  2HK2 A;  3PS1 A;  2UUA E;  1UDQ A;  1P42 A;  2DEI A;  3LTO A;  3QT5 A;  5DRR A;  4LFA E;  3GLL A;  1N0V C;  1N33 E;  3V2U C;  1NHJ A;  2PNZ B;  4OX9 I;  4LFB I;  4Z7Y A;  1PIE A;  5DRQ A;  4I3H A;  2UU9 I;  1ELO A;  2ZBK B;  4DPW A;  2IOP A;  4B3M I;  1USU A;  4U3D A;  4NXM I;  1HNZ I;  2VQF E;  3HJC A;  2NPF A;  2JE6 A;  2UUC I;  2NN6 A;  3GON A;  4DV3 I;  2CGE A;  4DUZ I;  5IT9 Q;  2CVE A;  4DR5 I;  5DNX A;  1MU5 A;  4JI7 E;  2EFG A;  5EKW A;  2VQE I;  3LPS A;  4DU7 A;  4AQY E;  2J65 A;  5L3J A;  2C37 A;  2F1D A;  4IZJ A;  4MU4 A;  4DV4 E;  1HR0 I;  4ED4 A;  1OYR A;  4X65 I;  3K85 A;  3Q6N A;  4YY3 E;  1FKA E;  4UTG A;  1H7U A;  1KVK A;  4FN5 A;  4B3T I;  1PKP A;  1XXE A;  5FLX C;  2BM1 A;  2NN6 C;  3QT6 A;  3J80 Q;  3PS2 A;  1A6F A;  4LF7 E;  4DXL A;  4JI8 E;  4LFC I;  1DAR A;  1R6L A;  4FW9 A;  3GME A;  4X64 E;  2PNM A;  4EMD A;  2UUA I;  1KIJ A;  4LFA I;  2PO2 A;  2PNL A;  1KTV A;  4FWG A;  2JE6 B;  5ELW A;  4LF9 E;  2NN6 B;  1N33 I;  4JI5 E;  2V8P A;  4U3B A;  4OX9 E;  2PO1 B;  5JEA E;  2UXB E;  2O1T A;  2GQ0 A;  3M6A A;  2Z5C A;  3CWV A;  4BA2 A;  4DR6 E;  3D4J A;  3UHM A;  1Z5A A;  1I94 E;  3WU3 A;  4IZJ B;  2HFS A;  2VQF I;  1FI4 A;  4J3D A;  2Z5B B;  4OO1 C;  4LOM A;  1FWK A;  2VES A;  3B78 A;  4WUC A;  1D6T A;  4DUY E;  1H73 A;  2JEB B;  3ZM7 A;  4JI3 I;  5FWK A;  4JI7 I;  1UDO A;  1Y60 A;  4GQU A;  2C39 A;  1UDN A;  2Q2E B;  3NZK A;  4OO1 B;  1KKH A;  4JI0 E;  5LL6 R;  3B4T A;  3PRY A;  1FNM A;  5IWA E;  4DV4 I;  1Z5C A;  3ZKD A;  5C0X D;  1XNQ E;  5EL9 A;  4USM A;  2AE8 A;  1N32 E;  4YY3 I;  2E5L E;  1Z0W A;  5GMD A;  3DD6 A;  2HKE A;  4LF7 I;  2WNR A;  4JI8 I;  3V5R A;  4IFD A;  2BA0 G;  2A2C A;  2IOQ A;  4KQV A;  3J81 C;  3PS3 A;  5LMU E;  2CZ9 A;  3PYE A;  4LF5 E;  5K36 A;  3M7N G;  4IFD D;  1E3H A;  2BR2 A;  1S4E A;  4HXZ A;  1MG7 A;  3U1Y A;  4JI4 E;  2V2Q A;  3HKM A;  4DR2 E;  2NN6 E;  2BM0 A;  3H4L A;  4LF9 I;  4JI5 I;  5K36 D;  3WU6 A;  4JI1 E;  4DV6 E;  2PO0 A;  2UXB I;  4IZK A;  3T1Y E;  4P7A A;  4MDT A;  4DR6 I;  2R3V A;  1I94 I;  1XHK A;  1YH8 A;  1U2R A;  4WUB A;  4HYM A;  3P76 A;  5A2Q Q;  3Q6M A;  1J5E E;  1Z59 A;  1HNW E;  2O1V A;  4B3R E;  5GME A;  3WU5 A;  4DUY I;  4GKJ E;  1Z0B A;  4JI3 E;  2BA1 G;  5DRP A;  2GO3 A;  4DV0 E; 
#chains in the Genus database with same CATH homology
 3P3G A;  4ISA A;  4LCG A;  4LCH A;  2J65 A;  2VES A;  4MQY A;  4MDT A;  4U3D A;  4IS9 A;  2GO4 A;  3P3E A;  4U3B A;  4FW3 A;  4FW5 A;  1YHC A;  1YH8 A;  4FW6 A;  3PS1 A;  3P76 A;  1P42 A;  3PS3 A;  2IER A;  1XXE A;  5DRR A;  3NZK A;  4FW7 A;  4OKG A;  4OZE A;  3U1Y A;  3PS2 A;  2JT2 A;  3UHM A;  4LCF A;  4FW4 A;  2O3Z A;  5DRP A;  2IES A;  4J3D A;  2GO3 A;  5DRQ A;  3P3C A;  5DRO A; 
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similar chains in the Genus database (?% sequence similarity)
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similar chains in the pdb database (?% sequence similarity)

 
#similar chains in the Genus database (?% sequence similarity)
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#similar chains, but unknotted
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#similar chains in the pdb database (?% sequence similarity)
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