4MQYA

Crystal structure of the escherichia coli lpxc/lpc-138 complex
Total Genus 87
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The genus trace: a function that shows values of genus (vertical axis) for subchains spanned between the first residue, and all other residues (shown on horizontal axis). The number of the latter residue and the genus of a given subchain are shown interactively.

Total Genus
87
sequence length
302
structure length
302
Chain Sequence
b'MIKQRTLKRIVQATGVGLHTGKKVTLTLRPAPANTGVIYRRTDLNPPVDFPADAKSVRDTMLCTCLVNEHDVRISTVEHLNAALAGLGIDNIVIEVNAPEIPIMDGSAAPFVYLLLDAGIDELNCAKKFVRIKETVRVEDGDKWAEFKPYNGFSLDFTIDFNHPAIDSSNQRYAMNFSADAFMRQISRARTFGFMRDIEYLQSRGLCLGGSFDCAIVVDDYRVLNEDGLRFEDEFVRHKMLDAIGDLFMCGHNIIGAFTAYKSGHALNNKLLQAVLAKQEAWEYVTFQDDAELPLAFKAPSA'
Loading diagram...

The genus matrix. At position (x,y) a genus value for a subchain spanned between x’th and y’th residue is shown. Values of the genus are represented by color, according to the scale given on the right.

Structure visualization

After clicking on a point (x,y) in the genus matrix above, a subchain from x to y is shown in color.

Chord Diagram
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molecule keywords UDP-3-O-[3-hydroxymyristoyl] N-acetylglucosamine deacetylase
molecule tags Hydrolase/hydrolase inhibitor
source organism Escherichia coli
publication title Structural Basis of the Promiscuous Inhibitor Susceptibility of Escherichia coli LpxC.
pubmed doi rcsb
total genus 87
structure length 302
sequence length 302
ec nomenclature ec 3.5.1.108: UDP-3-O-acyl-N-acetylglucosamine deacetylase.
pdb deposition date 2013-09-17

pfam database annotations

chain Pfam Accession Code Pfam Family Identifier Pfam Description
A PF03331 LpxC UDP-3-O-acyl N-acetylglycosamine deacetylase
Image from the rcsb pdb (www.rcsb.org)
cath code
ClassArchitectureTopologyHomologyDomain
3.30.230.20 Alpha Beta 2-Layer Sandwich Ribosomal Protein S5; domain 2 lpxc deacetylase, domain 1 4mqyA01
3.30.1700.10 Alpha Beta 2-Layer Sandwich lpxc deacetylase, domain 1 lpxc deacetylase, domain 2 4mqyA02
5DRRA 3PS2A 4LCGA 5DROA 5DRQA 2VESA 1P42A 3P3CA 1YH8A 3PS1A 4FW4A 4FW5A 3PS3A 4J3DA 3P76A 4ISAA 4OKGA 4FW6A 2IERA 3U1YA 4FW7A 4LCFA 2GO4A 4LCHA 4U3DA 2IESA 3P3EA 4FW3A 3NZKA 2JT2A 4MQYA 3UHMA 4MDTA 4IS9A 2J65A 3P3GA 5DRPA 1YHCA 4U3BA 2O3ZA 1XXEA 2GO3A 4OZEA
chains in the Genus database with same CATH superfamily
1XNQI 2E5LI 5EKWA 2HKEA 1XNRE 4OX9E 3PYGA 3M7NG 1NHJA 5K36C 1XNQE 4Z7CA 1H73A 4X62I 1ZM3A 2NN6F 4LF8E 2UXBE 4YHHI 1QZRA 3ZZTA 3V5RA 3DD6A 4JI3I 4OZEA 5K36B 4K0KE 4MYTA 4OO1B 4DR6I 5DROA 3J81C 2E5LE 4LPFA 5GMEA 1PVGA 3PYEA 1OYSA 4IFDC 4R1FA 5FLXQ 4USMA 2JEAA 4FW3A 3NZKA 1B62A 3QT8A 2NN6E 5DNLA 1FKAE 4OO1F 4U3BA 4PRVA 4JI2I 4LF4E 4DR6E 1IBLE 1R6LA 4DPYA 2GQ0A 5DRRA 5JEAC 3PS2A 3M85D 5IT9Q 2O1VA 4IFDA 2VESA 4JI0E 2Z5EA 3WU5A 4JI7I 4DR7E 2BV3A 2UUBI 5ELWA 2X7IA 4J3DA 3HKMA 3ZM7A 4DV2I 1ZM4A 3J81Q 1Z0VA 4LF9I 2C38B 4B3TI 4EMDA 2JT2A 1USVA 4IS9A 3H4LA 4GKKE 4AQYE 5K36F 4DR2E 4JI0I 5A2QQ 3M7ND 2WP8A 5LMNI 1DARA 4FWGA 4DPXA 2HHHE 2DEJA 1MG7A 4WUCA 3PS3A 4NBQA 4DV1E 4I3HA 1VISA 2NN6B 2CG9A 3L7ZB 4JI8I 2GO4A 4JI3E 4KHPE 2BA1G 4DU8A 2BM1A 5C0XB 4UTGA 4JI8E 4DU7A 3UHMA 1KTVA 4DXLA 1KIJA 2GS8A 1UEKA 4IFDD 4HACB 2NN6C 1N0VC 3T1HI 3LNUA 4AQYI 4LOMA 1HNWE 3GONA 4KQVA 3B78A 1ELOA 4DV5E 4LF8I 2UXBI 5C0XA 2WW4A 4U3DA 2PNZA 5L3JA 4B3RE 3J80Q 3B8HA 4FW9A 5IWAI 2OI2A 3K17A 2Z5BA 3M85G 1XXEA 2GO3A 4OX9I 4DR3I 4LFAI 1BKNA 3J7AM 3L7ZA 4MU4A 3T1HE 3K1JA 4DR5E 3V2UC 4DR2I 1YH8A 5C0XD 2PO0A 4NXME 1U2RA 4OKGA 4LFAE 2Z5CB 3ZZ0A 2F1DA 1FJGE 4HY1A 3U1YA 2Q2EB 1FJGI 4DV0E 4HACA 3GMEA 4ZVIA 1Z0TA 3HULA 3P3EA 4X64E 2HFSA 5FLXC 3QT7A 4X66I 2E1RA 4LFBI 2C39B 1XMQI 4MU1A 3Q6NA 1N36I 4FN5A 3CDIA 4DV3I 4UT4A 1HR0I 1NZ0A 1WUUA 2A2CA 5IWAE 3ZKDA 5EL9A 2IOPA 2PNMA 3Q1QA 1FWLA 3CDJA 2ZBKB 1Z0BA 3D4JA 2PO0B 3M6AA 1Y4SA 1IBMI 4JI7E 4M1KA 4BA1B 2HFUA 4JI1I 1D6TA 1Y4UA 4AIDA 3OTOI 1IBKI 4LCGA 1IBLI 2BA1D 5JEAE 3F0NA 4FW4A 1RREA 4X64I 1N32E 4IZKA 1Y6ZA 3ZZUA 4GKKI 4JI5I 4QNJA 4KHPI 4LF7I 3ZKBA 3JAMC 3GLLA 4OO1D 3U1KA 4BA1A 2ZM6I 1VI7A 4FWDA 3JAMQ 4IZJA 1IBKE 2NPFA 4X65I 4DV3E 5K36D 2O1WA 2JEAB 1KKHA 5K36A 1UDSA 4JI4E 1N36E 3K85A 1H7UA 1FWKA 2PO2B 1J5EI 4HXZA 5JEAF 4LFBE 1RHYA 2V2QA 4IFDB 4Z7YA 1N33I 4AIMA 2C37B 4BA2A 3WU6A 4DV2E 2V34A 5IT9C 4DR1E 5DRPA 1OYRA 1YHCA 4DV0I 4N3OA 4JI4I 4DR5I 4DV6E 1N32I 4X65E 2NN6D 2DEIA 4LF6I 2JEBB 1H72C 1N0UA 1UDNA 1Z0GA 4DV6I 4USKA 3J80C 4JYAE 2C37A 1MU5A 4BA2B 1K47A 4DR1I 4RPFA 1H7SA 4JYAI 5LL6R 1N34I 1PKPA 4P52A 3WU3A 4MDTA 1OYPA 3P3GA 1A6FA 2NN6A 3PYFA 4B3SI 3LPSA 1B63A 2UUBE 1Z5BA 3B82A 5LMUI 2F4VE 1HR0E 2PO1A 4IFDE 1NHHA 1Z59A 3PS1A 5C0XC 2R42A 2CVEA 1Z0CA 5LMUE 2UUCI 2ZM6E 2EFGA 4DV4I 4JI6E 4XTJA 1MX0A 5JEAD 2IESA 1S16A 2WNRB 2ZITA 4JI6I 4DV5I 1ZM2A 3B4TA 4OO1A 2PNLA 4DV7E 2V2VA 3KRNA 1J5EE 2UUCE 1E3HA 1KVKA 2O3ZA 4PRXA 2AE8A 4B3ME 2BA0G 3T1YE 1HNZI 4B3SE 4OO1E 4ED4A 4X66E 3OTOE 2Z5BB 3T1YI 1HNZE 3QT6A 1XMQE 4DR3E 5FWKA 4FWHA 2VQEE 2UXDE 2JE6A 4QNKA 2WNRA 3PRYA 2UXCI 2HK2A 3GCMA 2BM0A 1FNMA 2C38A 1I94I 4URLA 4LF7E 4DV7I 1Z5CA 1E3PA 5GMDA 4DV1I 2R3VA 4MU3A 2BA0D 2PO1B 4DUZE 4JG4A 4NXNE 1N33E 2UU9I 4WUBA 4DR7I 3PYDA 2BR2A 2A2DA 5BR8E 4K0KI 1IBME 5DNXA 5BR8I 1XNRI 4FW5A 1HNWI 4B3RI 3P76A 4MYUA 4DR4E 1N34E 5C0XE 4JI2E 1Z0WA 4HYMA 2WP8B 4P7AA 4DV4E 5JEAA 1I94E 2F4VI 2JE6B 2HKJA 2HHHI 4GKJE 4DUYI 4NXMI 4LF4I 4NXNI 4DPTA 4LF6E 1P42A 4LFCE 3P3CA 2IOQA 2PNZB 1Y5YA 4ISAA 4GKJI 3LTOA 2IERA 1S4EA 4IZJD 1Z0EA 1Z5AA 2CGEA 2UXDI 3J7AG 2LJPA 5K36E 2BR2B 4MQYA 4PU9A 2V2ZA 4DPWA 4DR4I 5C0XF 1XMOE 4LF5E 4LF9E 4DPUA 1EI1A 2VQFE 3P06A 4LFCI 1PIEA 1ZXNA 1EA6A 2V8PA 3CWVA 1USUA 1FI4A 4B3MI 2J7KA 2Z5CA 2VQFI 5JEAB 1ZM9A 4YHHE 4GQUA 1Y60A 4DUZI 4WUDA 1H74A 4FW7A 4B3TE 4YY3E 4IZKB 1XMOI 3WU4A 2HK3A 3HJCA 4JV5I 1OJ4A 1R6MA 2O1UA 5A2QC 2C39A 2J65A 1UDQA 2AJ4A 2PO2A 1NHIA 2CZ9A 4IZJB 4OO1C 1RR9A 2VQEI 2O1TA 4MU0A 2UUAE 5DRQA 4LF5I 1HNXE 2UU9E 4JI1E 1HK7A 2UUAI 4JI5E 2X36A 4FW6A 3H1CA 4IFDF 4LCFA 3Q6MA 2VF3A 4LCHA 1UDOA 2UXCE 2JEBA 3QT5A 4YY3I 4AM3A 4X62E 4JV5E 1ZXMA 1XHKA 2XEXA 5LMNE 4DUYE 1HNXI
chains in the Genus database with same CATH topology
5DRRA 3PS2A 4LCGA 5DROA 5DRQA 2VESA 1P42A 3P3CA 1YH8A 3PS1A 4FW4A 4FW5A 3PS3A 4J3DA 3P76A 4ISAA 4OKGA 4FW6A 2IERA 3U1YA 4FW7A 4LCFA 2GO4A 4LCHA 4U3DA 2IESA 3P3EA 4FW3A 3NZKA 2JT2A 4MQYA 3UHMA 4MDTA 4IS9A 2J65A 3P3GA 5DRPA 1YHCA 4U3BA 2O3ZA 1XXEA 2GO3A 4OZEA
chains in the Genus database with same CATH homology


 
#chains in the Genus database with same CATH superfamily
 5DRR A;  3PS2 A;  4LCG A;  5DRO A;  5DRQ A;  2VES A;  1P42 A;  3P3C A;  1YH8 A;  3PS1 A;  4FW4 A;  4FW5 A;  3PS3 A;  4J3D A;  3P76 A;  4ISA A;  4OKG A;  4FW6 A;  2IER A;  3U1Y A;  4FW7 A;  4LCF A;  2GO4 A;  4LCH A;  4U3D A;  2IES A;  3P3E A;  4FW3 A;  3NZK A;  2JT2 A;  4MQY A;  3UHM A;  4MDT A;  4IS9 A;  2J65 A;  3P3G A;  5DRP A;  1YHC A;  4U3B A;  2O3Z A;  1XXE A;  2GO3 A;  4OZE A; 
#chains in the Genus database with same CATH topology
 1XNQ I;  2E5L I;  5EKW A;  2HKE A;  1XNR E;  4OX9 E;  3PYG A;  3M7N G;  1NHJ A;  5K36 C;  1XNQ E;  4Z7C A;  1H73 A;  4X62 I;  1ZM3 A;  2NN6 F;  4LF8 E;  2UXB E;  4YHH I;  1QZR A;  3ZZT A;  3V5R A;  3DD6 A;  4JI3 I;  4OZE A;  5K36 B;  4K0K E;  4MYT A;  4OO1 B;  4DR6 I;  5DRO A;  3J81 C;  2E5L E;  4LPF A;  5GME A;  1PVG A;  3PYE A;  1OYS A;  4IFD C;  4R1F A;  5FLX Q;  4USM A;  2JEA A;  4FW3 A;  3NZK A;  1B62 A;  3QT8 A;  2NN6 E;  5DNL A;  1FKA E;  4OO1 F;  4U3B A;  4PRV A;  4JI2 I;  4LF4 E;  4DR6 E;  1IBL E;  1R6L A;  4DPY A;  2GQ0 A;  5DRR A;  5JEA C;  3PS2 A;  3M85 D;  5IT9 Q;  2O1V A;  4IFD A;  2VES A;  4JI0 E;  2Z5E A;  3WU5 A;  4JI7 I;  4DR7 E;  2BV3 A;  2UUB I;  5ELW A;  2X7I A;  4J3D A;  3HKM A;  3ZM7 A;  4DV2 I;  1ZM4 A;  3J81 Q;  1Z0V A;  4LF9 I;  2C38 B;  4B3T I;  4EMD A;  2JT2 A;  1USV A;  4IS9 A;  3H4L A;  4GKK E;  4AQY E;  5K36 F;  4DR2 E;  4JI0 I;  5A2Q Q;  3M7N D;  2WP8 A;  5LMN I;  1DAR A;  4FWG A;  4DPX A;  2HHH E;  2DEJ A;  1MG7 A;  4WUC A;  3PS3 A;  4NBQ A;  4DV1 E;  4I3H A;  1VIS A;  2NN6 B;  2CG9 A;  3L7Z B;  4JI8 I;  2GO4 A;  4JI3 E;  4KHP E;  2BA1 G;  4DU8 A;  2BM1 A;  5C0X B;  4UTG A;  4JI8 E;  4DU7 A;  3UHM A;  1KTV A;  4DXL A;  1KIJ A;  2GS8 A;  1UEK A;  4IFD D;  4HAC B;  2NN6 C;  1N0V C;  3T1H I;  3LNU A;  4AQY I;  4LOM A;  1HNW E;  3GON A;  4KQV A;  3B78 A;  1ELO A;  4DV5 E;  4LF8 I;  2UXB I;  5C0X A;  2WW4 A;  4U3D A;  2PNZ A;  5L3J A;  4B3R E;  3J80 Q;  3B8H A;  4FW9 A;  5IWA I;  2OI2 A;  3K17 A;  2Z5B A;  3M85 G;  1XXE A;  2GO3 A;  4OX9 I;  4DR3 I;  4LFA I;  1BKN A;  3J7A M;  3L7Z A;  4MU4 A;  3T1H E;  3K1J A;  4DR5 E;  3V2U C;  4DR2 I;  1YH8 A;  5C0X D;  2PO0 A;  4NXM E;  1U2R A;  4OKG A;  4LFA E;  2Z5C B;  3ZZ0 A;  2F1D A;  1FJG E;  4HY1 A;  3U1Y A;  2Q2E B;  1FJG I;  4DV0 E;  4HAC A;  3GME A;  4ZVI A;  1Z0T A;  3HUL A;  3P3E A;  4X64 E;  2HFS A;  5FLX C;  3QT7 A;  4X66 I;  2E1R A;  4LFB I;  2C39 B;  1XMQ I;  4MU1 A;  3Q6N A;  1N36 I;  4FN5 A;  3CDI A;  4DV3 I;  4UT4 A;  1HR0 I;  1NZ0 A;  1WUU A;  2A2C A;  5IWA E;  3ZKD A;  5EL9 A;  2IOP A;  2PNM A;  3Q1Q A;  1FWL A;  3CDJ A;  2ZBK B;  1Z0B A;  3D4J A;  2PO0 B;  3M6A A;  1Y4S A;  1IBM I;  4JI7 E;  4M1K A;  4BA1 B;  2HFU A;  4JI1 I;  1D6T A;  1Y4U A;  4AID A;  3OTO I;  1IBK I;  4LCG A;  1IBL I;  2BA1 D;  5JEA E;  3F0N A;  4FW4 A;  1RRE A;  4X64 I;  1N32 E;  4IZK A;  1Y6Z A;  3ZZU A;  4GKK I;  4JI5 I;  4QNJ A;  4KHP I;  4LF7 I;  3ZKB A;  3JAM C;  3GLL A;  4OO1 D;  3U1K A;  4BA1 A;  2ZM6 I;  1VI7 A;  4FWD A;  3JAM Q;  4IZJ A;  1IBK E;  2NPF A;  4X65 I;  4DV3 E;  5K36 D;  2O1W A;  2JEA B;  1KKH A;  5K36 A;  1UDS A;  4JI4 E;  1N36 E;  3K85 A;  1H7U A;  1FWK A;  2PO2 B;  1J5E I;  4HXZ A;  5JEA F;  4LFB E;  1RHY A;  2V2Q A;  4IFD B;  4Z7Y A;  1N33 I;  4AIM A;  2C37 B;  4BA2 A;  3WU6 A;  4DV2 E;  2V34 A;  5IT9 C;  4DR1 E;  5DRP A;  1OYR A;  1YHC A;  4DV0 I;  4N3O A;  4JI4 I;  4DR5 I;  4DV6 E;  1N32 I;  4X65 E;  2NN6 D;  2DEI A;  4LF6 I;  2JEB B;  1H72 C;  1N0U A;  1UDN A;  1Z0G A;  4DV6 I;  4USK A;  3J80 C;  4JYA E;  2C37 A;  1MU5 A;  4BA2 B;  1K47 A;  4DR1 I;  4RPF A;  1H7S A;  4JYA I;  5LL6 R;  1N34 I;  1PKP A;  4P52 A;  3WU3 A;  4MDT A;  1OYP A;  3P3G A;  1A6F A;  2NN6 A;  3PYF A;  4B3S I;  3LPS A;  1B63 A;  2UUB E;  1Z5B A;  3B82 A;  5LMU I;  2F4V E;  1HR0 E;  2PO1 A;  4IFD E;  1NHH A;  1Z59 A;  3PS1 A;  5C0X C;  2R42 A;  2CVE A;  1Z0C A;  5LMU E;  2UUC I;  2ZM6 E;  2EFG A;  4DV4 I;  4JI6 E;  4XTJ A;  1MX0 A;  5JEA D;  2IES A;  1S16 A;  2WNR B;  2ZIT A;  4JI6 I;  4DV5 I;  1ZM2 A;  3B4T A;  4OO1 A;  2PNL A;  4DV7 E;  2V2V A;  3KRN A;  1J5E E;  2UUC E;  1E3H A;  1KVK A;  2O3Z A;  4PRX A;  2AE8 A;  4B3M E;  2BA0 G;  3T1Y E;  1HNZ I;  4B3S E;  4OO1 E;  4ED4 A;  4X66 E;  3OTO E;  2Z5B B;  3T1Y I;  1HNZ E;  3QT6 A;  1XMQ E;  4DR3 E;  5FWK A;  4FWH A;  2VQE E;  2UXD E;  2JE6 A;  4QNK A;  2WNR A;  3PRY A;  2UXC I;  2HK2 A;  3GCM A;  2BM0 A;  1FNM A;  2C38 A;  1I94 I;  4URL A;  4LF7 E;  4DV7 I;  1Z5C A;  1E3P A;  5GMD A;  4DV1 I;  2R3V A;  4MU3 A;  2BA0 D;  2PO1 B;  4DUZ E;  4JG4 A;  4NXN E;  1N33 E;  2UU9 I;  4WUB A;  4DR7 I;  3PYD A;  2BR2 A;  2A2D A;  5BR8 E;  4K0K I;  1IBM E;  5DNX A;  5BR8 I;  1XNR I;  4FW5 A;  1HNW I;  4B3R I;  3P76 A;  4MYU A;  4DR4 E;  1N34 E;  5C0X E;  4JI2 E;  1Z0W A;  4HYM A;  2WP8 B;  4P7A A;  4DV4 E;  5JEA A;  1I94 E;  2F4V I;  2JE6 B;  2HKJ A;  2HHH I;  4GKJ E;  4DUY I;  4NXM I;  4LF4 I;  4NXN I;  4DPT A;  4LF6 E;  1P42 A;  4LFC E;  3P3C A;  2IOQ A;  2PNZ B;  1Y5Y A;  4ISA A;  4GKJ I;  3LTO A;  2IER A;  1S4E A;  4IZJ D;  1Z0E A;  1Z5A A;  2CGE A;  2UXD I;  3J7A G;  2LJP A;  5K36 E;  2BR2 B;  4MQY A;  4PU9 A;  2V2Z A;  4DPW A;  4DR4 I;  5C0X F;  1XMO E;  4LF5 E;  4LF9 E;  4DPU A;  1EI1 A;  2VQF E;  3P06 A;  4LFC I;  1PIE A;  1ZXN A;  1EA6 A;  2V8P A;  3CWV A;  1USU A;  1FI4 A;  4B3M I;  2J7K A;  2Z5C A;  2VQF I;  5JEA B;  1ZM9 A;  4YHH E;  4GQU A;  1Y60 A;  4DUZ I;  4WUD A;  1H74 A;  4FW7 A;  4B3T E;  4YY3 E;  4IZK B;  1XMO I;  3WU4 A;  2HK3 A;  3HJC A;  4JV5 I;  1OJ4 A;  1R6M A;  2O1U A;  5A2Q C;  2C39 A;  2J65 A;  1UDQ A;  2AJ4 A;  2PO2 A;  1NHI A;  2CZ9 A;  4IZJ B;  4OO1 C;  1RR9 A;  2VQE I;  2O1T A;  4MU0 A;  2UUA E;  5DRQ A;  4LF5 I;  1HNX E;  2UU9 E;  4JI1 E;  1HK7 A;  2UUA I;  4JI5 E;  2X36 A;  4FW6 A;  3H1C A;  4IFD F;  4LCF A;  3Q6M A;  2VF3 A;  4LCH A;  1UDO A;  2UXC E;  2JEB A;  3QT5 A;  4YY3 I;  4AM3 A;  4X62 E;  4JV5 E;  1ZXM A;  1XHK A;  2XEX A;  5LMN E;  4DUY E;  1HNX I; 
#chains in the Genus database with same CATH homology
 5DRR A;  3PS2 A;  4LCG A;  5DRO A;  5DRQ A;  2VES A;  1P42 A;  3P3C A;  1YH8 A;  3PS1 A;  4FW4 A;  4FW5 A;  3PS3 A;  4J3D A;  3P76 A;  4ISA A;  4OKG A;  4FW6 A;  2IER A;  3U1Y A;  4FW7 A;  4LCF A;  2GO4 A;  4LCH A;  4U3D A;  2IES A;  3P3E A;  4FW3 A;  3NZK A;  2JT2 A;  4MQY A;  3UHM A;  4MDT A;  4IS9 A;  2J65 A;  3P3G A;  5DRP A;  1YHC A;  4U3B A;  2O3Z A;  1XXE A;  2GO3 A;  4OZE A; 
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similar chains in the Genus database (?% sequence similarity)
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similar chains in the pdb database (?% sequence similarity)

 
#similar chains in the Genus database (?% sequence similarity)
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#similar chains, but unknotted
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#similar chains in the pdb database (?% sequence similarity)
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