4LCFA

Crystal structure of the pseudomonas aeruginosa lpxc/lpc-014 complex
Total Genus 95
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The genus trace: a function that shows values of genus (vertical axis) for subchains spanned between the first residue, and all other residues (shown on horizontal axis). The number of the latter residue and the genus of a given subchain are shown interactively.

Total Genus
95
sequence length
298
structure length
298
Chain Sequence
b'MIKQRTLKNIIRATGVGLHSGEKVYLTLKPAPVDTGIVFSRTDLDPVVEIPARAENVGETTMSTTLVKGDVKVDTVEHLLSAMAGLGIDNAYVELSASEVPIMDGSAGPFVFLIQSAGLQEQEAAKKFIRIKREVSVEEGDKRAVFVPFDGFKVSFEIDFDHPVFRGRTQQASVDFSSTSFVKEVSRARTFGFMRDIEYLRSQNLALGGSVENAIVVDENRVLNEDGLRYEDEFVKHKILDAIGDLYLLGNSLIGEFRGFKSGHALNNQLLRTLIADKDAWEVVTFEDARTAPISYMR'
Loading diagram...

The genus matrix. At position (x,y) a genus value for a subchain spanned between x’th and y’th residue is shown. Values of the genus are represented by color, according to the scale given on the right.

Structure visualization

After clicking on a point (x,y) in the genus matrix above, a subchain from x to y is shown in color.

Chord Diagram
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molecule keywords UDP-3-O-[3-hydroxymyristoyl] N-acetylglucosamine deacetylase
molecule tags Hydrolase/antibiotic
source organism Pseudomonas aeruginosa
publication title Synthesis, Structure, and Antibiotic Activity of Aryl-Substituted LpxC Inhibitors.
pubmed doi rcsb
total genus 95
structure length 298
sequence length 298
ec nomenclature ec 3.5.1.108: UDP-3-O-acyl-N-acetylglucosamine deacetylase.
pdb deposition date 2013-06-21

pfam database annotations

chain Pfam Accession Code Pfam Family Identifier Pfam Description
A PF03331 LpxC UDP-3-O-acyl N-acetylglycosamine deacetylase
Image from the rcsb pdb (www.rcsb.org)
cath code
ClassArchitectureTopologyHomologyDomain
3.30.230.20 Alpha Beta 2-Layer Sandwich Ribosomal Protein S5; domain 2 lpxc deacetylase, domain 1 4lcfA01
3.30.1700.10 Alpha Beta 2-Layer Sandwich lpxc deacetylase, domain 1 lpxc deacetylase, domain 2 4lcfA02
3PS1A 3P3GA 4J3DA 5DRPA 2IESA 4OZEA 4U3BA 4OKGA 4FW3A 4FW6A 4IS9A 3PS3A 4LCHA 2JT2A 2O3ZA 1XXEA 2GO4A 4LCGA 2J65A 3P3EA 4FW7A 4LCFA 4U3DA 5DRRA 5DROA 3UHMA 1YHCA 4MDTA 4MQYA 2IERA 3NZKA 3PS2A 5DRQA 2VESA 2GO3A 4FW4A 4FW5A 1P42A 3P3CA 3P76A 3U1YA 1YH8A 4ISAA
chains in the Genus database with same CATH superfamily
2ZM6I 4YHHE 4IFDF 1PVGA 4X66E 2ZM6E 3QT6A 2BA1D 4JI5I 4IFDB 2HHHE 1HK7A 4MU4A 3OTOE 4JI4E 4FWGA 4LOMA 4Z7CA 2F1DA 2DEIA 1Y4UA 2VF3A 1Z5AA 1N34E 3V2UC 1Z0EA 4JI4I 3ZZ0A 4LF6I 1XNRE 2IERA 3ZZTA 2OI2A 2F4VI 5LL6R 1BKNA 4JYAI 1VISA 3L7ZA 1B62A 1P42A 1HNWI 1N33E 5IWAE 4DR7E 5IT9Q 2JEAA 2IESA 3LNUA 4OX9I 4JG4A 5FLXQ 4AM3A 4FW6A 4WUDA 2EFGA 4X62E 5IWAI 4DR7I 3H1CA 1NHIA 4DPTA 5K36A 4FW9A 4BA2A 4IZJD 2VQEI 1FJGE 4OO1F 2VQEE 1UDNA 4LF4I 4DV6I 4DR2E 4DR5I 4NBQA 5GMDA 1PKPA 4P7AA 4ISAA 4PRVA 1U2RA 1K47A 2UUAI 4DV7I 2HKEA 3T1HI 2CGEA 1Z0GA 3PS3A 4LF8E 2IOPA 3WU6A 2JE6B 1MU5A 5JEAD 4FW7A 4DV1E 4YY3I 5C0XE 4OO1B 1WUUA 3PRYA 1N34I 5EL9A 3JAMQ 4NXNE 2JEBA 3KRNA 5LMUI 4JI0E 1Y5YA 2XEXA 1NHJA 1XNQI 2PO0A 1S4EA 1NHHA 2CZ9A 3M85G 3T1YE 1N36I 1ZXNA 1XNRI 3Q1QA 1HNZI 4USMA 1UDOA 2AJ4A 4JI1I 1XNQE 5FWKA 4X65I 3PYDA 1FWLA 2IOQA 1IBLI 2BA0G 3GONA 3P3EA 4DV7E 1N0UA 3OTOI 4DXLA 2NN6A 4LFCI 4DUYE 4LF7I 1MX0A 2PO1A 4GKJI 2Q2EB 2V2VA 1H72C 4MDTA 3M7ND 2PNZB 1UDQA 4IZKB 4LF6E 1KIJA 1QZRA 1ZXMA 1N32I 4JI0I 3Q6NA 2E1RA 4JI3E 1Y6ZA 1RREA 4JI8I 4DV6E 4DR2I 4DR5E 4B3MI 3UHMA 4MYTA 2UU9E 1Z0VA 1N33I 2WNRA 4WUCA 2NN6F 3GCMA 1J5EE 4QNJA 3WU5A 5JEAF 1USUA 1YH8A 4LCGA 4XTJA 2C39B 4PU9A 1Z0BA 4DV1I 4FW3A 4DR1E 5C0XC 2BR2A 4IS9A 5C0XB 3H4LA 1I94E 4GKKI 2BM0A 4U3DA 2WP8A 1OJ4A 2V2ZA 1Z5CA 3ZKBA 3F0NA 2CVEA 1HNXE 2Z5CB 1PIEA 4B3ME 3P06A 3P76A 4AIMA 2PO0B 5DRPA 1OYRA 2Z5EA 4PRXA 2UXBE 3V5RA 2JEBB 4M1KA 2BM1A 4DV0E 1H7SA 4MU1A 1KVKA 2HFUA 4DV4E 4DU7A 1ZM2A 2AE8A 4DU8A 5BR8I 2JE6A 4DR1I 1YHCA 1Y4SA 4B3TI 3L7ZB 2Z5CA 2BA1G 4JI8E 4P52A 5BR8E 1HNWE 3M7NG 4JI3I 1FWKA 3PYFA 3B8HA 3P3CA 4B3SI 2X36A 3PYGA 1HNZE 2UU9I 1XMQE 1OYSA 4DUZE 1UEKA 4B3SE 2C39A 1KTVA 1USVA 1Z0CA 1FKAE 4IFDA 2PNMA 4AIDA 5LMNE 3J7AG 4BA2B 4DUYI 4X62I 5DRRA 4LFCE 2V8PA 3J81Q 2UUBI 4KQVA 3B4TA 3PS2A 3U1KA 2C37B 2O1TA 4MU3A 2UXCI 1S16A 5GMEA 2O1UA 4JI7E 1HR0E 4X64E 2HFSA 2JEAB 4GKKE 3K1JA 5C0XD 1ZM3A 1Z5BA 3HJCA 5JEAE 2DEJA 4HYMA 1VI7A 2GQ0A 3HULA 1ELOA 4NXMI 2WP8B 2O1VA 5EKWA 4LF5E 4NXME 4DPYA 4DR3I 4JI2I 1D6TA 3PYEA 4OX9E 5C0XA 1Z0WA 3NZKA 2BR2B 3J81C 5LMNI 3CWVA 2NN6B 1J5EI 2HK3A 3WU4A 4K0KI 4DV5E 3QT5A 4Z7YA 2NN6D 4U3BA 4B3TE 2WNRB 4R1FA 5K36E 2V34A 4DV3I 1EA6A 4JI6I 2LJPA 4USKA 1HNXI 4JYAE 4DUZI 4DPUA 1IBMI 3GLLA 5FLXC 4YHHI 4LF7E 4MQYA 1IBME 2NPFA 5DRQA 3B78A 4DV3E 2O1WA 1Z0TA 1KKHA 1XHKA 4B3RE 4AQYE 4IZKA 2PNZA 1H7UA 4J3DA 1E3HA 2PO1B 1E3PA 5IT9C 4N3OA 4X66I 4DR6E 2GS8A 4X64I 4I3HA 3WU3A 2ZBKB 5C0XF 2E5LI 1H73A 2C38A 4LCFA 2V2QA 4LFBI 2GO3A 4JI2E 1MG7A 2R3VA 5JEAB 5K36B 3K85A 1XMQI 4IZJA 5K36D 4LFAE 1HR0I 4OO1A 2A2CA 2A2DA 1NZ0A 2PO2B 2GO4A 2UXDI 3ZKDA 5DNXA 4DR4E 2NN6E 2UUBE 3K17A 5DROA 1UDSA 4DV5I 3HKMA 2VESA 4DV2I 1XMOE 4HXZA 3LPSA 1FNMA 4LF8I 4FWDA 3U1YA 2NN6C 2VQFI 3T1HE 4LCHA 4DV2E 3PS1A 3B82A 3CDIA 1B63A 4OKGA 3CDJA 4FN5A 4JV5I 4AQYI 1OYPA 4JV5E 4JI5E 5A2QC 3ZZUA 4YY3E 1N36E 3J7AM 1IBKI 2X7IA 1Y60A 4URLA 3Q6MA 4JI6E 4DV4I 5ELWA 4DPXA 3DD6A 5L3JA 1IBKE 4LF9E 2HKJA 4UT4A 3P3GA 5A2QQ 2R42A 4KHPE 4RPFA 1A6FA 4MYUA 4UTGA 4IFDD 4OO1D 4X65E 4OO1E 4IFDE 4DR6I 4DV0I 3LTOA 2PO2A 1FJGI 1EI1A 2PNLA 4GKJE 3D4JA 4HACA 4IZJB 4FW5A 2UXCE 1N32E 3M85D 1Z59A 2HK2A 1RHYA 4ED4A 4LFBE 5K36C 4OZEA 4OO1C 5DNLA 2Z5BA 4DR4I 5JEAA 2E5LE 1DARA 3JAMC 2JT2A 2J7KA 4JI7I 2O3ZA 3QT7A 1XMOI 3QT8A 2C38B 2UXBI 4KHPI 4HACB 3J80C 2UUAE 2ZITA 4GQUA 4EMDA 1ZM4A 4DR3E 2UXDE 4BA1A 2WW4A 4LFAI 4HY1A 4LPFA 1N0VC 2UUCE 1ZM9A 4IFDC 3M6AA 4LF4E 1R6LA 1RR9A 2CG9A 4BA1B 1H74A 4QNKA 3T1YI 4DPWA 4K0KE 3GMEA 2UUCI 4FWHA 3ZM7A 4NXNI 2BV3A 4WUBA 1I94I 5LMUE 1XXEA 2F4VE 2J65A 4ZVIA 4LF9I 5JEAC 3J80Q 1IBLE 1R6MA 4LF5I 1FI4A 2C37A 2BA0D 2Z5BB 5K36F 4MU0A 4FW4A 2VQFE 4B3RI 4JI1E 2HHHI
chains in the Genus database with same CATH topology
3PS1A 3P3GA 4J3DA 5DRPA 2IESA 4OZEA 4U3BA 4OKGA 4FW3A 4FW6A 4IS9A 3PS3A 4LCHA 2JT2A 2O3ZA 1XXEA 2GO4A 4LCGA 2J65A 3P3EA 4FW7A 4LCFA 4U3DA 5DRRA 5DROA 3UHMA 1YHCA 4MDTA 4MQYA 2IERA 3NZKA 3PS2A 5DRQA 2VESA 2GO3A 4FW4A 4FW5A 1P42A 3P3CA 3P76A 3U1YA 1YH8A 4ISAA
chains in the Genus database with same CATH homology


 
#chains in the Genus database with same CATH superfamily
 3PS1 A;  3P3G A;  4J3D A;  5DRP A;  2IES A;  4OZE A;  4U3B A;  4OKG A;  4FW3 A;  4FW6 A;  4IS9 A;  3PS3 A;  4LCH A;  2JT2 A;  2O3Z A;  1XXE A;  2GO4 A;  4LCG A;  2J65 A;  3P3E A;  4FW7 A;  4LCF A;  4U3D A;  5DRR A;  5DRO A;  3UHM A;  1YHC A;  4MDT A;  4MQY A;  2IER A;  3NZK A;  3PS2 A;  5DRQ A;  2VES A;  2GO3 A;  4FW4 A;  4FW5 A;  1P42 A;  3P3C A;  3P76 A;  3U1Y A;  1YH8 A;  4ISA A; 
#chains in the Genus database with same CATH topology
 2ZM6 I;  4YHH E;  4IFD F;  1PVG A;  4X66 E;  2ZM6 E;  3QT6 A;  2BA1 D;  4JI5 I;  4IFD B;  2HHH E;  1HK7 A;  4MU4 A;  3OTO E;  4JI4 E;  4FWG A;  4LOM A;  4Z7C A;  2F1D A;  2DEI A;  1Y4U A;  2VF3 A;  1Z5A A;  1N34 E;  3V2U C;  1Z0E A;  4JI4 I;  3ZZ0 A;  4LF6 I;  1XNR E;  2IER A;  3ZZT A;  2OI2 A;  2F4V I;  5LL6 R;  1BKN A;  4JYA I;  1VIS A;  3L7Z A;  1B62 A;  1P42 A;  1HNW I;  1N33 E;  5IWA E;  4DR7 E;  5IT9 Q;  2JEA A;  2IES A;  3LNU A;  4OX9 I;  4JG4 A;  5FLX Q;  4AM3 A;  4FW6 A;  4WUD A;  2EFG A;  4X62 E;  5IWA I;  4DR7 I;  3H1C A;  1NHI A;  4DPT A;  5K36 A;  4FW9 A;  4BA2 A;  4IZJ D;  2VQE I;  1FJG E;  4OO1 F;  2VQE E;  1UDN A;  4LF4 I;  4DV6 I;  4DR2 E;  4DR5 I;  4NBQ A;  5GMD A;  1PKP A;  4P7A A;  4ISA A;  4PRV A;  1U2R A;  1K47 A;  2UUA I;  4DV7 I;  2HKE A;  3T1H I;  2CGE A;  1Z0G A;  3PS3 A;  4LF8 E;  2IOP A;  3WU6 A;  2JE6 B;  1MU5 A;  5JEA D;  4FW7 A;  4DV1 E;  4YY3 I;  5C0X E;  4OO1 B;  1WUU A;  3PRY A;  1N34 I;  5EL9 A;  3JAM Q;  4NXN E;  2JEB A;  3KRN A;  5LMU I;  4JI0 E;  1Y5Y A;  2XEX A;  1NHJ A;  1XNQ I;  2PO0 A;  1S4E A;  1NHH A;  2CZ9 A;  3M85 G;  3T1Y E;  1N36 I;  1ZXN A;  1XNR I;  3Q1Q A;  1HNZ I;  4USM A;  1UDO A;  2AJ4 A;  4JI1 I;  1XNQ E;  5FWK A;  4X65 I;  3PYD A;  1FWL A;  2IOQ A;  1IBL I;  2BA0 G;  3GON A;  3P3E A;  4DV7 E;  1N0U A;  3OTO I;  4DXL A;  2NN6 A;  4LFC I;  4DUY E;  4LF7 I;  1MX0 A;  2PO1 A;  4GKJ I;  2Q2E B;  2V2V A;  1H72 C;  4MDT A;  3M7N D;  2PNZ B;  1UDQ A;  4IZK B;  4LF6 E;  1KIJ A;  1QZR A;  1ZXM A;  1N32 I;  4JI0 I;  3Q6N A;  2E1R A;  4JI3 E;  1Y6Z A;  1RRE A;  4JI8 I;  4DV6 E;  4DR2 I;  4DR5 E;  4B3M I;  3UHM A;  4MYT A;  2UU9 E;  1Z0V A;  1N33 I;  2WNR A;  4WUC A;  2NN6 F;  3GCM A;  1J5E E;  4QNJ A;  3WU5 A;  5JEA F;  1USU A;  1YH8 A;  4LCG A;  4XTJ A;  2C39 B;  4PU9 A;  1Z0B A;  4DV1 I;  4FW3 A;  4DR1 E;  5C0X C;  2BR2 A;  4IS9 A;  5C0X B;  3H4L A;  1I94 E;  4GKK I;  2BM0 A;  4U3D A;  2WP8 A;  1OJ4 A;  2V2Z A;  1Z5C A;  3ZKB A;  3F0N A;  2CVE A;  1HNX E;  2Z5C B;  1PIE A;  4B3M E;  3P06 A;  3P76 A;  4AIM A;  2PO0 B;  5DRP A;  1OYR A;  2Z5E A;  4PRX A;  2UXB E;  3V5R A;  2JEB B;  4M1K A;  2BM1 A;  4DV0 E;  1H7S A;  4MU1 A;  1KVK A;  2HFU A;  4DV4 E;  4DU7 A;  1ZM2 A;  2AE8 A;  4DU8 A;  5BR8 I;  2JE6 A;  4DR1 I;  1YHC A;  1Y4S A;  4B3T I;  3L7Z B;  2Z5C A;  2BA1 G;  4JI8 E;  4P52 A;  5BR8 E;  1HNW E;  3M7N G;  4JI3 I;  1FWK A;  3PYF A;  3B8H A;  3P3C A;  4B3S I;  2X36 A;  3PYG A;  1HNZ E;  2UU9 I;  1XMQ E;  1OYS A;  4DUZ E;  1UEK A;  4B3S E;  2C39 A;  1KTV A;  1USV A;  1Z0C A;  1FKA E;  4IFD A;  2PNM A;  4AID A;  5LMN E;  3J7A G;  4BA2 B;  4DUY I;  4X62 I;  5DRR A;  4LFC E;  2V8P A;  3J81 Q;  2UUB I;  4KQV A;  3B4T A;  3PS2 A;  3U1K A;  2C37 B;  2O1T A;  4MU3 A;  2UXC I;  1S16 A;  5GME A;  2O1U A;  4JI7 E;  1HR0 E;  4X64 E;  2HFS A;  2JEA B;  4GKK E;  3K1J A;  5C0X D;  1ZM3 A;  1Z5B A;  3HJC A;  5JEA E;  2DEJ A;  4HYM A;  1VI7 A;  2GQ0 A;  3HUL A;  1ELO A;  4NXM I;  2WP8 B;  2O1V A;  5EKW A;  4LF5 E;  4NXM E;  4DPY A;  4DR3 I;  4JI2 I;  1D6T A;  3PYE A;  4OX9 E;  5C0X A;  1Z0W A;  3NZK A;  2BR2 B;  3J81 C;  5LMN I;  3CWV A;  2NN6 B;  1J5E I;  2HK3 A;  3WU4 A;  4K0K I;  4DV5 E;  3QT5 A;  4Z7Y A;  2NN6 D;  4U3B A;  4B3T E;  2WNR B;  4R1F A;  5K36 E;  2V34 A;  4DV3 I;  1EA6 A;  4JI6 I;  2LJP A;  4USK A;  1HNX I;  4JYA E;  4DUZ I;  4DPU A;  1IBM I;  3GLL A;  5FLX C;  4YHH I;  4LF7 E;  4MQY A;  1IBM E;  2NPF A;  5DRQ A;  3B78 A;  4DV3 E;  2O1W A;  1Z0T A;  1KKH A;  1XHK A;  4B3R E;  4AQY E;  4IZK A;  2PNZ A;  1H7U A;  4J3D A;  1E3H A;  2PO1 B;  1E3P A;  5IT9 C;  4N3O A;  4X66 I;  4DR6 E;  2GS8 A;  4X64 I;  4I3H A;  3WU3 A;  2ZBK B;  5C0X F;  2E5L I;  1H73 A;  2C38 A;  4LCF A;  2V2Q A;  4LFB I;  2GO3 A;  4JI2 E;  1MG7 A;  2R3V A;  5JEA B;  5K36 B;  3K85 A;  1XMQ I;  4IZJ A;  5K36 D;  4LFA E;  1HR0 I;  4OO1 A;  2A2C A;  2A2D A;  1NZ0 A;  2PO2 B;  2GO4 A;  2UXD I;  3ZKD A;  5DNX A;  4DR4 E;  2NN6 E;  2UUB E;  3K17 A;  5DRO A;  1UDS A;  4DV5 I;  3HKM A;  2VES A;  4DV2 I;  1XMO E;  4HXZ A;  3LPS A;  1FNM A;  4LF8 I;  4FWD A;  3U1Y A;  2NN6 C;  2VQF I;  3T1H E;  4LCH A;  4DV2 E;  3PS1 A;  3B82 A;  3CDI A;  1B63 A;  4OKG A;  3CDJ A;  4FN5 A;  4JV5 I;  4AQY I;  1OYP A;  4JV5 E;  4JI5 E;  5A2Q C;  3ZZU A;  4YY3 E;  1N36 E;  3J7A M;  1IBK I;  2X7I A;  1Y60 A;  4URL A;  3Q6M A;  4JI6 E;  4DV4 I;  5ELW A;  4DPX A;  3DD6 A;  5L3J A;  1IBK E;  4LF9 E;  2HKJ A;  4UT4 A;  3P3G A;  5A2Q Q;  2R42 A;  4KHP E;  4RPF A;  1A6F A;  4MYU A;  4UTG A;  4IFD D;  4OO1 D;  4X65 E;  4OO1 E;  4IFD E;  4DR6 I;  4DV0 I;  3LTO A;  2PO2 A;  1FJG I;  1EI1 A;  2PNL A;  4GKJ E;  3D4J A;  4HAC A;  4IZJ B;  4FW5 A;  2UXC E;  1N32 E;  3M85 D;  1Z59 A;  2HK2 A;  1RHY A;  4ED4 A;  4LFB E;  5K36 C;  4OZE A;  4OO1 C;  5DNL A;  2Z5B A;  4DR4 I;  5JEA A;  2E5L E;  1DAR A;  3JAM C;  2JT2 A;  2J7K A;  4JI7 I;  2O3Z A;  3QT7 A;  1XMO I;  3QT8 A;  2C38 B;  2UXB I;  4KHP I;  4HAC B;  3J80 C;  2UUA E;  2ZIT A;  4GQU A;  4EMD A;  1ZM4 A;  4DR3 E;  2UXD E;  4BA1 A;  2WW4 A;  4LFA I;  4HY1 A;  4LPF A;  1N0V C;  2UUC E;  1ZM9 A;  4IFD C;  3M6A A;  4LF4 E;  1R6L A;  1RR9 A;  2CG9 A;  4BA1 B;  1H74 A;  4QNK A;  3T1Y I;  4DPW A;  4K0K E;  3GME A;  2UUC I;  4FWH A;  3ZM7 A;  4NXN I;  2BV3 A;  4WUB A;  1I94 I;  5LMU E;  1XXE A;  2F4V E;  2J65 A;  4ZVI A;  4LF9 I;  5JEA C;  3J80 Q;  1IBL E;  1R6M A;  4LF5 I;  1FI4 A;  2C37 A;  2BA0 D;  2Z5B B;  5K36 F;  4MU0 A;  4FW4 A;  2VQF E;  4B3R I;  4JI1 E;  2HHH I; 
#chains in the Genus database with same CATH homology
 3PS1 A;  3P3G A;  4J3D A;  5DRP A;  2IES A;  4OZE A;  4U3B A;  4OKG A;  4FW3 A;  4FW6 A;  4IS9 A;  3PS3 A;  4LCH A;  2JT2 A;  2O3Z A;  1XXE A;  2GO4 A;  4LCG A;  2J65 A;  3P3E A;  4FW7 A;  4LCF A;  4U3D A;  5DRR A;  5DRO A;  3UHM A;  1YHC A;  4MDT A;  4MQY A;  2IER A;  3NZK A;  3PS2 A;  5DRQ A;  2VES A;  2GO3 A;  4FW4 A;  4FW5 A;  1P42 A;  3P3C A;  3P76 A;  3U1Y A;  1YH8 A;  4ISA A; 
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similar chains in the Genus database (?% sequence similarity)
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similar chains in the pdb database (?% sequence similarity)

 
#similar chains in the Genus database (?% sequence similarity)
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#similar chains, but unknotted
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#similar chains in the pdb database (?% sequence similarity)
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